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FOC | Family List | FOC Vol. 19 | Acanthaceae

29. Barleria Linnaeus, Sp. Pl. 2: 636. 1753.

假杜鹃属 jia du juan shu

Authors: Jia-qi Hu & Thomas F. Daniel

Barleria cristata

Credit: Harvard University Herbaria

Shrubs, subshrubs, or perennial herbs, with cystoliths, usually spiny. Leaves opposite, petiolate. Inflorescences axillary cymes (sometimes secund or reduced to dense clusters), terminal spikes, or flowers solitary; bracts present or absent; bracteoles 2, sometimes becoming spinose. Calyx deeply 4-lobed; outer 2 lobes larger [anterior lobe sometimes 2-cleft]; inner 2 lobes smaller. Corolla funnel-shaped, usually large; limb 5-lobed (usually with upper lip appearing 4-lobed); lobes subequal, quincuncial in bud. Stamens [2 or]4 and didynamous, one pair usually greatly reduced and/or sterile; anthers 2-thecous; thecae ± equal in size, equally inserted, lacking basal appendages; staminodes 1 or 3, sometimes bearing very small anthers. Ovary with 2 ovules per locule; stigma 2-cleft or entire. Capsule substipitate or lacking a distinct basal stalk, subellipsoid to ovoid, 2-4-seeded; retinacula present. Seeds discoid, pubescent with appressed hygroscopic trichomes.

About 80-120 species: tropical regions of Africa and Asia, one species in tropical America; four species (one endemic) in China.

Barleria procumbens Loureiro (Fl. Cochinch. 2: 377. 1790) is not treated here because no material was available for study. Barleria lupulina Lindley is only cultivated in China and not treated here.

1 Margin of calyx lobes (at least outer lobes) serrate or spiny   (2)
+ Margin of calyx lobes entire (apex spine-tipped in B. prionitis)   (3)
2 (1) Flowers in dense, shortly pedunculate cymes from leaf axils; bracteoles linear to linear-lanceolate, 2.4-6.5 cm.   1 B. cristata
+ Flowers in axillary and terminal dense spikes; bracteoles elliptic, ca. 1.2 cm.   3 B. strigosa
3 (1) Plants with spines in at least some leaf axils; calyx lobes apically spinose; corolla yellowish to orange.   2 B. prionitis
+ Plants without spines; calyx lobes not apically spinose; corolla not yellowish to orange   (4)
4 (3) Flowers in dense spikes; calyx purple; narrow basal portion of corolla tube ca. 3 × as long as throat; bracteoles ca. 1/2 as long as calyx; seeds 4 per capsule.   3 B. strigosa
+ Flowers solitary in leaf axils; calyx green; narrow basal portion of corolla tube shorter than throat; bracteoles equaling or longer than calyx; seeds 2 per capsule.   4 B. integrisepala

Lower Taxa


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