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33. Chiritopsis W. T. Wang, Bull. Bot. Res., Harbin. 1(3): 21. 1981.

小花苣苔属 xiao hua ju tai shu

Herbs, perennial, epipetric or terrestrial, rhizomatous, stemless. Leaves few to many, basal; leaf blade puberulent to pilose, base cuneate to cordate. Inflorescences lax or rarely dense, axillary, few- to many-flowered cymes; bracts 2, opposite. Calyx actinomorphic, 5-sect from base; segments equal. Corolla white, yellowish, or purplish, zygomorphic, inside puberulent to glabrous; tube nearly tubular, not swollen, much longer than limb, 3-5 mm in diam.; limb indistinctly to distinctly 2-lipped; adaxial lip 2-lobed, slightly to 2-3 X shorter than abaxial lip; abaxial lip 3(or 4)-lobed, lobes equal or central lobe longer, apex rounded. Stamens 2, adnate to abaxial side of corolla tube near base or above middle, included; anthers dorsifixed, connate by adaxial surfaces, thecae divaricate, confluent at apex, dehiscing longitudinally; connective not projecting; staminodes (absent or)1 or 3, adnate to adaxial or adaxial and abaxial sides of corolla tube. Disc ringlike or 2-lobed. Ovary ovoid, 1-loculed; placentas 2, parietal, projecting inward, 2-cleft. Stigma 1, abaxial, obtrapeziform, rarely linear, undivided to 2-lobed. Capsule straight in relation to pedicel, ovoid to narrowly ellipsoid, equalling to shorter than calyx, dehiscing loculicidally to base; valves 2, straight, not twisted. Seeds unappendaged.

* Nine species: endemic to China.


1 Leaf blade margin pinnately to palmately lobed; staminodes 1 or 3, less often 2 or absent.   (2)
+ Leaf blade margin denticulate, dentate, crenate, or nearly entire; staminodes 2 or absent.   (4)
       
2 (1) Leaf blade ovate to rhombic, base broadly cuneate, margin pinnatifid   7 Chiritopsis bipinnatifida
+ Leaf blade ovate to orbicular or reniform, base cordate, margin palmately or nearly palmately lobed.   (3)
       
3 (2) Corolla ca. 7.8 mm; petiole, peduncle, pedicel, and calyx white puberulent to pilose; calyx segments 3-4 X 0.4-0.7 mm; stamens glabrous; disc 2-lobed, eglandular; staminode 1   8 Chiritopsis lobulata
+ Corolla 11-13 mm; petiole, peduncle, pedicel, and calyx glandular puberulent or pubescent and purple glandular; calyx segments 3-8 X 1.3-2 mm; stamens sparsely pubescent; disc ringlike, glandular; staminodes 2 or 3, seldom absent   9 Chiritopsis glandulosa
       
4 (1) Leaf blade orbicular-ovate to nearly orbicular; corolla purplish; stigma linear, 1-1.5 mm, undivided   1 Chiritopsis mollifolia
+ Leaf blade elliptic or rhombic to ovate or reniform, rarely nearly orbicular; corolla white or yellowish; stigma obtrapeziform, 0.3-0.9 mm, 2-lobed.   (5)
       
5 (4) Leaf blade base rounded to cordate; peduncle 5-9 cm; corolla 10-14 mm, inside puberulent in 2 longitudinal lines on or below abaxial lip; stamens puberulent or sparsely so.   (6)
+ Leaf blade base cuneate to broadly cuneate, seldom to nearly cordate; peduncle 8-9 cm; corolla 7.2-12 mm, inside glabrous or sparsely puberulent below abaxial lip; stamens glabrous.   (7)
       
6 (5) Leaf blade 4-8.5 X 3-9 cm, puberulent, margin repand-crenate to obtusely dentate; petiole 5.5-16.5 cm; bracts 2-3 X ca. 0.3 mm; pedicel 4-18 mm; corolla ca. 1.4 cm; stamens adnate to corolla ca. 3 mm above base, staminodes 2   5 Chiritopsis cordifolia
+ Leaf blade 3-4 X 2.2-3.6 cm, pubescent and pilose, margin remotely denticulate to entire; petiole 3-7 cm; bracts 5.5-7 X 1.8-2 mm; pedicel 1.5-3.5 mm; corolla ca. 1 cm; stamens adnate to corolla 1-1.5 mm above base, staminodes absent   6 Chiritopsis subulata
       
7 (5) Cymes dense; pedicels 0.2-2 mm; corolla distinctly 2-lipped, adaxial lip ca. 0.8 mm, abaxial lip ca. 2.2 mm, inside sparsely puberulent below abaxial lip; disc ringlike, ca. 0.5 mm; style ca. 2.8 mm, puberulent   4 Chiritopsis confertiflora
+ Cymes lax; pedicels 3-25 mm; corolla indistinctly 2-lipped, adaxial lip 2-3 mm, abaxial lip ca. 2 mm, inside glabrous; disc 2-lobed, 1-1.5 mm; style 5-7 mm, glabrous or nearly so.   (8)
       
8 (7) Leaf blade puberulent to villous, base cuneate to broadly cuneate; lateral veins conspicuous; pedicel 3-8 mm; corolla white, ca. 8 mm   2 Chiritopsis repanda
+ Leaf blade sparsely puberulent, base broadly cuneate to nearly cordate; lateral veins inconspicuous; pedicel 8-25 mm; corolla yellowish, ca. 12 mm   3 Chiritopsis xiuningensis

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