1 |
Inflorescences racemose; seeds with scalelike appendages at both ends; flowers sometimes replaced by propagules |
|
56 Titanotrichum |
+ |
Flowers solitary or inflorescences cymose (including umbel-like or paniculate cymes), sometimes racemelike and 1-sided; seeds either not appendiculate or with subulate or hairlike appendages at 1 or both ends (unknown in Calcareoboea. Dayaoshania, Gyrogyne. Lagarosolen. Paraisometrum,and Thamnocharis); flowers never replaced by propagules. |
|
(2) |
|
|
|
|
2 (1) |
Corolla actinomorphic, 4- or 5-lobed; stamens 4 or 5, staminodes absent; seeds unappendaged (unknown in Thamnocharis). |
|
(3) |
+ |
Corolla zygomorphic, usually distinctly 2-lipped or rarely actinomorphic; fertile stamens (1 or)2 or 4, staminodes absent or 1-3; seeds appendaged or unappendaged. |
|
(6) |
|
|
|
|
3 (2) |
Corolla suburceolate, tube longer than limb; stamens included, anther thecae divergent, confluent apically; capsule 4-valved |
|
3 Tengia |
+ |
Corolla rotate or campanulate, tube 1/3-1 X limb; stamens equalling corolla or exserted, anther thecae parallel, not confluent; capsule 2-valved. |
|
(4) |
|
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|
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4 (3) |
Anthers connate into a tube surrounding style, connectives with long apical projections; disc absent; inflorescences umbel-like cymes; leaves 1 to few |
|
4 Conandron |
+ |
Anthers free, connectives without projections; disc ringlike; inflorescences cymes; leaves few to many. |
|
(5) |
|
|
|
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5 (4) |
Bracts 2, opposite; corolla purple to blue, deeply 4- or 5-lobed, tube rotate; stamens adnate to corolla near base, anthers basifixed; stigma 1, nearly truncate or punctiform; leaf blade base cuneate |
|
1 Thamnocharis |
+ |
Bracts 6-9 to numerous, crowded, involucrate; corolla white, 4- or 5-lobed, tube campanulate; stamens adnate to corolla tube near or above middle, anthers dorsifixed; stigmas 2, equal, oblate to 2-lipped; leaf blade base truncate to cordate |
|
2 Bournea |
|
|
|
|
6 (2) |
Fruit indehiscent berries, fleshy to leathery; plants woody herbs or shrubs. |
|
(7) |
+ |
Fruit dehiscent capsule; plants herbaceous to shrubby. |
|
(8) |
|
|
|
|
7 (6) |
Calyx divided from base; corolla campanulate-tubular; stamens 4 |
|
49 Rhynchotechum |
+ |
Calyx divided only from above middle (in China); corolla funnelform-tubular; stamens 2 (in China) |
|
50 Cyrtandra |
|
|
|
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8 (6) |
Calyx plicate between lobes; corolla tube campanulate, spurred or saccate at base; stamens 4; ovary depressed globose or subglobose. |
|
(9) |
+ |
Calyx not plicate; corolla tube funnelform to tubular, cylindric, salverform, urceolate, or campanulate, base usually not swollen, rarely saccate, sometimes gibbous abaxially; stamens (1 or)2 or 4; ovary linear to oblong or ovoid, rarely subglobose or globose. |
|
(10) |
|
|
|
|
9 (8) |
Leaves slightly unequal in a pair; adaxial corolla lip ca. 1/2 as long as abaxial lip; anthers free, thecae not confluent |
|
51 Gyrogyne |
+ |
Leaves strongly unequal in a pair, smaller much reduced; adaxial corolla lip as long as abaxial lip; anthers coherent laterally to form a shallow cone, thecae confluent at apex |
|
52 Stauranthera |
|
|
|
|
10 (8) |
Capsule ovoid to depressed globose, shorter than calyx; ovary ovoid to depressed globose; inflorescences cymose, sympodial, or scorpioid, racemose; bract apparently absent or 1; stem leaves 1 to few. |
|
(11) |
+ |
Capsule linear to ovoid, much surpassing calyx, rarely shorter than calyx, then leaves all basal (fruit unknown in Paraisometrum); ovary usually linear to oblong, rarely ellipsoid, linear-lanceolate, conical, or ovoid; flowers solitary or inflorescences cymose including paniculate or umbel-like cymes; bracts 2, rarely apparently absent, 1, or 3; leaves basal or on stem, few to many. |
|
(13) |
|
|
|
|
11 (10) |
Leaves opposite, strongly unequal in a pair, smaller much reduced and sessile; inflorescences apparently without bracts; calyx with many longitudinal secretory canals; staminode 1; ovary 2-loculed; placenta 1, axile |
|
53 Whytockia |
+ |
Leaves alternate, or lowest stem leaf only unpaired, adaxial ones usually opposite and equal or nearly so in a pair; bract 1; calyx without secretory canals; staminodes 2, 3, or absent; ovary 1-loculed; placentas 2, parietal. |
|
(12) |
|
|
|
|
12 (11) |
Leaves few, all alternate; inflorescences lax racemes appearing scorpioid; stamens 2 and abaxial, or 4; capsule loculicidal |
|
54 Rhynchoglossum |
+ |
Leaves 1 to few, lowest leaf unpaired, those near apex usually opposite; inflorescences scorpioid cymes, dense; stamens 2, adaxial; capsule circumscissile |
|
55 Epithema |
|
|
|
|
13 (10) |
Fertile stamens 4. |
|
(14) |
+ |
Fertile stamens 2. |
|
(37) |
|
|
|
|
14 (13) |
Calyx zygomorphic, 2-lipped, adaxial lip undivided, abaxial lip 4-lobed; disc tubular; capsule narrowly ellipsoid |
|
15 Cathayanthe |
+ |
Calyx actinomorphic, 5-lobed to 5-sect from base; disc ringlike, rarely cupular or inconspicuous; capsule oblong to linear, rarely conical to ovoid. |
|
(15) |
|
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|
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15 (14) |
Plants stemless. |
|
(16) |
+ |
Plants with a distinct stem. |
|
(27) |
|
|
|
|
16 (15) |
Adaxial corolla lip 4-lobed, over 2 X longer than abaxial lip, abaxial lip undivided |
|
8 Paraisometrum |
+ |
Adaxial corolla lip 2-lobed, equalling or much shorter than abaxial lip, abaxial lip 3-lobed or 3-sect. |
|
(17) |
|
|
|
|
17 (16) |
Stigma 1. |
|
(18) |
+ |
Stigmas 2. |
|
(21) |
|
|
|
|
18 (17) |
Abaxial corolla lip densely bearded inside; bracts apparently absent; corolla tube tubular; anthers coherent in pairs, thecae divaricate |
|
13 Corallodiscus |
+ |
Corolla glabrous to puberulent inside; bracts 2, rarely absent or several; corolla tube campanulate to broadly tubular, sometimes oblique, sometimes saccate at base, throat some- times constricted; anthers free or all 4 coherent together, thecae parallel or subparallel. |
|
(19) |
|
|
|
|
19 (18) |
Anthers of all 4 stamens coherent together; capsule oblique in relation to pedicel; adaxial corolla lip distinctly shorter than abaxial |
|
16 Beccarinda |
+ |
Anthers free; capsule straight in relation to pedicel; adaxial corolla lip slightly shorter than to as long as abaxial. |
|
(20) |
|
|
|
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20 (19) |
Ovary oblong; anther thecae parallel, not confluent, seldom horseshoe-shaped, then thecae divaricate and confluent at apex; corolla tube 1-4 X longer than limb; leaf blade base cuneate to subcordate |
|
5 Oreocharis |
+ |
Ovary ovoid or conical; anther thecae subparallel, confluent at apex; corolla tube slightly shorter than limb; leaf blade base cuneate to attenuate |
|
18 Boeica |
|
|
|
|
21 (17) |
Adaxial corolla lip emarginate to nearly entire. |
|
(22) |
+ |
Adaxial corolla lip 2-lobed to 2-sect from base. |
|
(23) |
|
|
|
|
22 (21) |
Corolla tube subcampanulate to funnelform-tubular, not swollen; stamens adnate to corolla near base, all or only abaxial ones exserted, anthers free, thecae confluent at apex |
|
6 Tremacron |
+ |
Corolla tube narrowly funnelform to cylindric, slightly gibbous abaxially; stamens adnate to corolla tube above or rarely below middle, usually included, rarely exserted, anthers coherent in pairs, thecae not confluent |
|
9 Ancylostemon |
|
|
|
|
23 (21) |
Ovary 2-loculed, only adaxial locule fertile; placenta 1, axile; capsule oblique in relation to pedicel; cymes 1-3-flowered |
|
11 Briggsiopsis |
+ |
Ovary 1-loculed; placentas 2, parietal; capsule straight in relation to pedicel; cymes 1- to many flowered. |
|
(24) |
|
|
|
|
24 (23) |
Corolla tube narrowly campanulate, gibbous abaxially, (0.7-)1-2(-2.6) cm in diam., abaxial lip 3-lobed |
|
10 Briggsia |
+ |
Corolla tube cylindric to funnelform, broadly tubular, campanulate, urceolate, or salverform, not swollen, slightly gibbous abaxially, or slightly ampliate basally, 0.2-1(-1.2) cm in diam., abaxial lip 3-sect, rarely 3-lobed. |
|
(25) |
|
|
|
|
25 (24) |
Anthers dehiscing longitudinally from arcuate slits; corolla zygomorphic, rarely actinomorphic, tube campanulate, cylindric, or rarely urceolate or salverform, not swollen |
|
7 Isometrum |
+ |
Anthers dehiscing longitudinally, rarely transversely; corolla zygomorphic, tube campanulate-cylindric to broadly tubular, sometimes ampliate basally, throat sometimes constricted or narrowly funnelform to cylindric, slightly gibbous abaxially. |
|
(26) |
|
|
|
|
26 (25) |
Corolla purple to blue or white or yellow to red, tube campanulate-cylindric to broadly tubular, sometimes ampliate basally, throat sometimes constricted, adaxial lip 2-lobed; anthers free; calyx 5-sect from base |
|
5 Oreocharis |
+ |
Corolla orange to yellow or white, rarely pink, tube narrowly funnelform to cylindric, slightly gibbous abaxially, adaxial lip 2-lobed, emarginate, or rarely undivided; anthers coherent in pairs; calyx 5-sect from base to 5-lobed from above middle |
|
9 Ancylostemon |
|
|
|
|
27 (15) |
Stigmas 2. |
|
(28) |
+ |
Stigma 1. |
|
(31) |
|
|
|
|
28 (27) |
Ovary 2-loculed, only adaxial locule fertile; placenta 1, axile; capsule oblique in relation to pedicel; corolla tube tubular-funnelform; cymes 1-3-flowered |
|
11 Briggsiopsis |
+ |
Ovary 1-loculed; placentas 2, parietal; corolla tube cylindric to broadly tubular or narrowly campanulate, sometimes ampliate above; capsule straight in relation to pedicel; cymes 1- to many flowered. |
|
(29) |
|
|
|
|
29 (28) |
Corolla tube cylindric basally, ampliate above, not swollen |
|
12 Didissandra |
+ |
Corolla tube narrowly campanulate to broadly tubular, gibbous abaxially. |
|
(30) |
|
|
|
|
30 (29) |
Seeds unappendaged |
|
10 Briggsia |
+ |
Seeds with 1 hairlike appendage at each end |
|
46 Loxostigma |
|
|
|
|
31 (27) |
Calyx zygomorphic, 2-lipped, adaxial lip undivided, abaxial lip 4-lobed; disc tubular; capsule splitting along 1 suture |
|
15 Cathayanthe |
+ |
Calyx actinomorphic, 5-sect from base or 5-lobed, segments equal or unequal; disc ringlike, inconspicuous or absent; capsule 2(or 4)-valved. |
|
(32) |
|
|
|
|
32 (31) |
Leaves 1(or 2), apical on stem; leaf blade base cordate to auriculate; anthers dehiscing from fissures or cracks; capsule slightly longer than calyx |
|
14 Platystemma |
+ |
Leaves few to many, opposite, rarely whorled, spread along stem or crowded near stem apex; leaf blade base cuneate to attenuate or rounded, rarely cordate; anthers dehiscing longitudinally, poricidally, or transversely; capsule much longer than calyx. |
|
(33) |
|
|
|
|
33 (32) |
Seeds with 1 subulate or hairlike appendage at each end, or with up to 50 appendages at hilar end; corolla tube narrowly tubular to funnelform-tubular or subtubular, longer or much longer than limb, 4-17 mm in diam.; ovary linear; stamens adnate to corolla tube near or above middle, anthers usually coherent in pairs. |
|
(34) |
+ |
Seeds unappendaged; corolla tube campanulate, shorter than to slightly longer than limb, 3-9 mm in diam.; ovary ovoid, ellipsoid, or conical; stamens adnate to corolla near or slightly above base, anthers free or all 4 coherent together. |
|
(35) |
|
|
|
|
34 (33) |
Corolla white to yellowish, tube subtubular, gibbous abaxially near limb, with 2 arcuate projections abaxially inside; stamens included, thecae confluent at apex; seeds with 1 subulate appendage at each end |
|
45 Anna |
+ |
Corolla red to orange, seldom greenish, yellow, or white, tube narrowly tubular to funnelform-tubular, often curved, not swollen, inside sparsely puberulent, sparsely glandular puberulent, glabrous, or with a hair ring; stamens usually exserted, thecae not confluent; seeds with 1 hairlike appendage at each end or up to 50 appendages at hilar end |
|
47 Aeschynanthus |
|
|
|
|
35 (33) |
All anthers coherent together; capsule oblique in relation to pedicel; leaf blade base cordate to cuneate |
|
16 Beccarinda |
+ |
Anthers free; capsule straight in relation to pedicel; leaf blade base cuneate to attenuate. |
|
(36) |
|
|
|
|
36 (35) |
Corolla yellowish, tube slightly longer than limb; anther thecae not confluent; ovary ellipsoid; cymes 3-5-flowered |
|
17 Leptobaea |
+ |
Corolla white to pink or blue to purple, tube slightly shorter than limb; anther thecae confluent at apex; ovary ovoid or conical; cymes 1- to many flowered |
|
18 Boeica |
|
|
|
|
37 (13) |
Plants with a distinct stem. |
|
(38) |
+ |
Plants stemless. |
|
(50) |
|
|
|
|
38 (37) |
Stigmas 2. |
|
(39) |
+ |
Stigma 1. |
|
(43) |
|
|
|
|
39 (38) |
Flowers solitary; capsule valves spirally twisted; corolla tube campanulate-tubular, adaxial lip slightly shorter than abaxial |
|
44 Rhabdothamnopsis |
+ |
Inflorescences cymes, sometime spikelike or racemelike, 2-10-flowered; capsule valves straight, not twisted; corolla tube slender funnelform, funnelform, or funnelform-tubular, adaxial lip much shorter than abaxial. |
|
(40) |
|
|
|
|
40 (39) |
Bracts forming a globose involucre; corolla tube much longer than limb; leaf blade base attenuate to broadly cuneate; stamens adnate to corolla tube below middle, anthers coherent by adaxial surfaces, thecae divaricate, confluent |
|
26 Hemiboeopsis |
+ |
Bracts not forming an involucre; corolla tube longer than limb; leaf blade base cuneate to rounded or cordate; stamens adnate to corolla tube near middle, anthers coherent at apex, thecae parallel, not confluent at apex. |
|
(41) |
|
|
|
|
41 (40) |
Stigmas equal, oblate or semiorbicular, undivided; calyx 5-sect from base; stamens adnate to corolla tube near mouth, slightly exserted |
|
21 Didymostigma |
+ |
Stigmas unequal, smaller 1 truncate or triangular, undivided, larger 1 narrowly obtrapeziform, emarginate; calyx 5-lobed; stamens adnate to corolla tube near middle, included. |
|
(42) |
|
|
|
|
42 (41) |
Ovary 1-loculed; placentas parietal; corolla abaxial lip lobes unequal |
|
22 Pseudochirita |
+ |
Ovary 2-loculed; placentas axile; corolla abaxial lip lobes subequal |
|
23 Allostigma |
|
|
|
|
43 (38) |
Subshrubs or rarely lianas, usually epiphytic, less frequently epipetric or terrestrial; seeds with 1 appendage at each end |
|
48 Lysionotus |
+ |
Herbs, rarely subshrubs (rarely shrubs outside China), always terrestrial or epipetric; seeds unappendaged. |
|
(44) |
|
|
|
|
44 (43) |
Anthers basifixed, thecae parallel, not confluent at apex; bracts 2, forming a globose to ovoid involucre. |
|
(45) |
+ |
Anthers dorsifixed, thecae divaricate, confluent at apex; bracts 2, not forming an involucre, rarely up to 7 and whorled, 1, or absent. |
|
(46) |
|
|
|
|
45 (44) |
Ovary 1-loculed; placenta parietal; capsule dehiscing on both sides; corolla inside glabrous; tube somewhat gibbous abaxially, adaxial lip inconspicuous, much shorter than abaxial lip |
|
24 Metabriggsia |
+ |
Ovary 2-loculed, only adaxial locule fertile; placenta axile; capsule dehiscing only adaxially; corolla inside usually with a ring of hairs; tube not swollen, adaxial lip shorter than to nearly as long as abaxial |
|
25 Hemiboea |
|
|
|
|
46 (44) |
Stamens adnate to abaxial side of corolla tube near or above middle; capsule valves straight, not twisted. |
|
(47) |
+ |
Stamens adnate to abaxial side of corolla tube near base; capsule valves usually at least slightly twisted. |
|
(48) |
|
|
|
|
47 (46) |
Stigma abaxial, 2-lobed to undivided |
|
32 Chirita |
+ |
Stigma terminal, undivided |
|
35 Didymocarpus |
|
|
|
|
48 (46) |
Anthers dehiscing transversely; disc cupular; stigma ligulate; calyx zygomorphic |
|
43 Trisepalum |
+ |
Anthers dehiscing longitudinally from straight or arcuate slits; disc inconspicuous or ringlike; stigma capitate, seldom subligulate; calyx actinomorphic, rarely slightly zygomorphic. |
|
(49) |
|
|
|
|
49 (48) |
Leaf blade adaxially woolly to glabrous, abaxially woolly, pannose, or velutinous, hairs interwoven, usually branched; corolla inside glabrous to glandular puberulent; tube longer than to occasionally nearly equalling limb |
|
40 Paraboea |
+ |
Leaf blade puberulent to woolly, hairs unbranched; corolla inside bearded, tube shorter than limb |
|
42 Ornithoboea |
|
|
|
|
50 (37) |
Corolla adaxial lip 4-lobed, abaxial lip undivided. |
|
(51) |
+ |
Corolla adaxial lip 2-lobed, rarely undivided, abaxial lip 3-lobed, rarely 4-lobed. |
|
(52) |
|
|
|
|
51 (50) |
Inflorescences dense umbel-like cymes; corolla tube 4-5 X longer than limb, adaxial lip longer than abaxial; stamens adnate to corolla tube above middle |
|
38 Calcareoboea |
+ |
Inflorescences lax cymes; corolla tube shorter than limb, adaxial lip slightly shorter than abaxial; stamens adnate to corolla tube near base |
|
39 Allocheilos |
|
|
|
|
52 (50) |
Anthers dehiscing transversely; stigma 1, ligulate; capsule valves spirally twisted |
|
43 Trisepalum |
+ |
Anthers dehiscing longitudinally; stigmas 1 or 2, if 1 then usually capitate, globose, disclike, truncate, lamelliform, oblate, or obtrapeziform, rarely linear or subligulate; capsule valves straight or twisted. |
|
(53) |
|
|
|
|
53 (52) |
Corolla adaxial lip undivided. |
|
(54) |
+ |
Corolla adaxial lip 2-lobed. |
|
(55) |
|
|
|
|
54 (53) |
Corolla tube tubular to funnelform-tubular, longer than limb; stamens exserted, anthers basifixed, free or coherent at apex, thecae parallel, not confluent; leaf blade base broadly cuneate to subrounded |
|
29 Deinocheilos |
+ |
Corolla tube broadly tubular, nearly equalling limb; stamens included, anthers dorsifixed, coherent by adaxial surfaces, thecae divaricate, confluent at apex; leaf blade base cordate |
|
36 Gyrocheilos |
|
|
|
|
55 (53) |
Anthers basifixed or if dorsifixed then stamens slightly exserted, thecae parallel. |
|
(56) |
+ |
Anthers dorsifixed or nearly so, stamens included, thecae divaricate. |
|
(58) |
|
|
|
|
56 (55) |
Anthers dorsifixed, stamens slightly exserted; placentas undivided; inflorescences monochasial cymes, 1- or 2-flowered |
|
20 Dayaoshania |
+ |
Anthers basifixed, stamens included; placentas projecting inward, 2-cleft; inflorescences cymes, sometimes umbel-like, 1- to many flowered. |
|
(57) |
|
|
|
|
57 (56) |
Corolla tube 1.5-4 X longer than limb, 3.5-17 mm in diam.; stamens adnate to adaxial side of corolla tube near middle, rarely near base; ovary and capsule linear |
|
19 Opithandra |
+ |
Corolla tube shorter than limb, 2-5 mm in diam.; stamens adnate to abaxial side of corolla tube near base; ovary conical to broadly ovoid; capsule narrowly ellipsoid to oblong to ovoid |
|
27 Petrocosmea |
|
|
|
|
58 (55) |
Anthers free, thecae not confluent; capsule nearly globose; leaf blade base rounded, peltate |
|
28 Metapetrocosmea |
+ |
Anthers coherent, thecae confluent at apex; capsule linear to narrowly ovoid, ellipsoid, or oblong; leaf blade base attenuate to cordate, rarely auriculate or auriculate-peltate. |
|
(59) |
|
|
|
|
59 (58) |
Stigmas 2; corolla lobe apex acute |
|
30 Lagarosolen |
+ |
Stigma 1; corolla lobe apex rounded, seldom obtuse or acute. |
|
(60) |
|
|
|
|
60 (59) |
Capsule valves spirally twisted or seldom straight, if straight leaf blade abaxially woolly, pannose, or velutinous, hairs dense, interwoven, usually branched. |
|
(61) |
+ |
Capsule valves straight, not twisted; leaf blade abaxially strigose or puberulent to villous, velutinous, or woolly. |
|
(62) |
|
|
|
|
61 (60) |
Capsule valves 4; leaf blade abaxially woolly, pannose, or velutinous, hairs very dense, interwoven, usually branched |
|
40 Paraboea |
+ |
Capsule valves 2; leaf blade villous to puberulent, hairs dense, not interwoven or branched |
|
41 Boea |
|
|
|
|
62 (60) |
Corolla tube slightly shorter than to nearly equalling limb, lobes acute |
|
37 Dolicholoma |
+ |
Corolla tube longer to much longer than limb, lobes rounded, obtuse, or acute. |
|
(63) |
|
|
|
|
63 (62) |
Ovary ovoid or narrowly ovoid; capsule narrowly ovoid to ellipsoid, equalling to shorter than calyx. |
|
(64) |
+ |
Ovary linear; capsule linear to very narrowly ovoid, much surpassing calyx. |
|
(65) |
|
|
|
|
64 (63) |
Corolla nearly salverform, tube cylindric, adaxial lip slightly shorter than abaxial |
|
31 Primulina |
+ |
Corolla tube nearly tubular, adaxial lip slightly to 2-3 X shorter than abaxial |
|
33 Chiritopsis |
|
|
|
|
65 (63) |
Corolla tube suburceolate-tubular, 1-2.5 mm in diam., limb indistinctly 2-lipped, adaxial lip slightly shorter than abaxial; lobe apex acute; stamens adnate to abaxial side of corolla tube slightly above base |
|
34 Petrocodon |
+ |
Corolla tube funnelform-tubular to campanulate or cylindric, 2-20 mm in diam., limb 2-lipped; adaxial lip slightly to 1.5-3.5 X shorter than abaxial; lobe apex rounded, rarely obtuse to acute; stamens adnate to abaxial side of corolla tube near or above middle. |
|
(66) |
|
|
|
|
66 (65) |
Stigma abaxial, 2-lobed to undivided |
|
32 Chirita |
+ |
Stigma terminal, undivided |
|
35 Didymocarpus |
|