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FOC | Family List | FOC Vol. 18 | Gesneriaceae

12. Didissandra C. B. Clarke in A. & C. de Candolle, Monogr. Phan. 5: 65. 1883.

漏斗苣苔属 lou dou ju tai shu

Raphiocarpus W. Y. Chun.

Herbs, rarely shrubs or subshrubs, perennial, epipetric or terrestrial, rhizomatous. Stems simple or branched. Leaves 2 to few, on stem, opposite, often crowded at stem apex, equal to strongly unequal in a pair; leaf blade puberulent to villous or glabrous, base cuneate to cordate. Inflorescences lax or dense, axillary, 1-10-flowered cymes; bracts 2 or apparently absent, opposite. Calyx actinomorphic, 5-sect from base to 5-lobed; segments equal. Corolla white to yellow, orange-red, purple, or blue-purple, zygomorphic, inside glabrous to sparsely glandular pubescent; tube narrowly cylindric to cylindric basally, ampliate above, not swollen, 4-5 X longer than limb, 0.6-1.5 cm in diam.; limb 2-lipped; adaxial lip 2-lobed, longer than to much shorter than abaxial lip; abaxial lip 3-lobed, lobes equal or subequal, apex rounded. Stamens 4, adnate to corolla tube near or above middle, included; anthers basifixed, coherent in pairs, thecae divergent, not confluent or confluent, dehiscing longitudinally; connective not projecting; staminode absent or 1, adnate to adaxial side of corolla tube. Disc ringlike. Ovary linear, 1-loculed; placentas 2, parietal, projecting inward, 2-cleft. Stigmas 2, equal or unequal, oblong to deltoid, undivided or 2-lobed. Capsule straight in relation to pedicel, linear to narrowly oblong, much longer than calyx, dehiscing loculicidally to base; valves 2, straight, not twisted. Seeds unappendaged.

About 31 species: China, Indonesia, Malaysia, Thailand, Vietnam; five species in China.

1 Leaves 1 pair, strongly unequal; petiole absent to 3 mm; calyx 5-lobed from below to above middle, tube at least 3 mm; stigmas unequal, adaxial undivided, oblong, ca. 2 mm, abaxial 2-lobed, obovate, ca. 3 mm   5 Didissandra sesquifolia
+ Leaves 2 or more pairs, equal to slightly unequal in each pair; petiole 0.3-6 cm; calyx 5-sect from base or near base, tube to 1 mm; stigmas equal, oblong to deltoid.   (2)
2 (1) Stem and peduncle glabrous to pubescent, glabrescent; calyx 2.5-6 mm; corolla yellow or white, often flushed purple, 1-4.9 cm.   (3)
+ Stem and peduncle villous to woolly, sometimes also puberulent or glabrescent; calyx 6-15 mm; corolla purple, purple-blue, or orange-red to yellow, 3-7 cm.   (4)
3 (2) Corolla white, often flushed purple, 1.2-1.5 cm; bracts 1-2 X ca. 0.5 mm; calyx 2.5-3 X ca. 1 mm; stems glabrous to very sparsely puberulent; leaf blade with both surfaces glabrous   1 Didissandra sinica
+ Corolla yellow, 3.5-4.9 cm; bracts ca. 3-6 X 1-3 mm; calyx 4-6 X 1-3 mm; stems pubescent to puberulent, glabrescent, leaf blade adaxially sparsely puberulent to strigose, abaxially glabrous to sparsely puberulent, veins sparsely pubescent to sparsely villous   2 Didissandra longipedunculata
4 (2) Cymes 5-10-flowered; peduncle 3-10(-12) cm; bracts deciduous, broadly ovate to orbicular, 15-25 X 15-30 mm, margin irregularly dentate to subentire; corolla pale-purple or rusty lilac to purple; ovary pubescent, style glandular puberulent   4 Didissandra begoniifolia
+ Cymes 1-3-flowered; peduncle 1.5-4.3 cm; bracts persistent, lanceolate to oblong or oblanceolate, 2-4 X ca. 1 mm, margin entire; corolla orange-red to yellow; ovary and style glabrous   3 Didissandra macrosiphon

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