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FOC | Family List | FOC Vol. 18 | Gesneriaceae

19. Opithandra B. L. Burtt, Baileya. 4: 162. 1956.

后蕊苣苔属 hou rui ju tai shu

Schistolobos W. T. Wang.

Herbs, perennial, epipetric or terrestrial, rhizomatous, stemless. Leaves few to many, basal; leaf blade puberulent to woolly or velutinous, base cuneate to cordate. Inflorescences often umbel-like, lax, axillary, 1- to many-flowered cymes; bracts 2, opposite. Calyx actinomorphic, 5-sect from base to deeply 5-lobed; segments equal. Corolla pink to purple, zygomorphic, inside glabrous to sparsely puberulent or glandular puberulent; tube tubular to funnelform, rarely nearly salverform or cylindric, not swollen, sometimes slightly gibbous abaxially, 1.5-4 X longer than limb, 3.5-17 mm in diam.; limb distinctly or indistinctly 2-lipped; adaxial lip 2-lobed, usually distinctly shorter than, rarely nearly as long as abaxial lip; abaxial lip 3-lobed, lobes equal or subequal, apex rounded to acute. Stamens 2, adnate to adaxial side of corolla tube near middle, rarely near corolla base, included; anthers basifixed, free or coherent at apex, thecae parallel, not confluent, dehiscing longitudinally from straight or arcuate slits; connective not projecting; staminodes (1-)3 or absent, adnate to adaxial, lower, or adaxial and abaxial sides of corolla tube. Disc ringlike or cupular. Ovary linear, 1-loculed; placentas 2, parietal, projecting inward, 2-cleft. Stigmas 2, equal, oblate, undivided, or 1, terminal, capitate, undivided or 2-lipped. Capsule straight in relation to pedicel, linear, much longer than calyx, dehiscing loculicidally to base; valves 2, straight, not twisted. Seeds unappendaged.

Ten species: China, Japan; nine species in China.


1 Corolla lobes lobed   9 Opithandra pumila
+ Corolla lobes entire.   (2)
       
2 (1) Anthers oblong or oblong ovoid with straight slits; corolla mouth 3-3.5 mm in diam.   (3)
+ Anthers globose-ovoid to reniform with arcuate slits; corolla mouth 5-17 mm in diam.   (6)
       
3 (2) Corolla tube nearly cylindric, puberulent outside, adaxial lip ca. 1 cm; leaf blade thick leathery, strongly bullate; pistil ca. 5.5 mm, stigma 1   4 Opithandra fargesii
+ Corolla tube tubular or nearly so, glabrous outside, adaxial lip 1-6 mm; leaf blade papery, nearly flat; pistil 6.5-14 mm; stigmas 1 or 2.   (4)
       
4 (3) Calyx segments ca. 2 mm; corolla to 1 cm, adaxial lip ca. 1 mm, abaxial lip ca. 2.8 mm; anthers coherent at apex; pistil exserted; stigma 1; lateral veins 4 or 5 on each side of midrib; bracts subulate   3 Opithandra acaulis
+ Calyx segments 3.5-5 mm; corolla at least 1.6 cm, adaxial lip 4-6 mm, abaxial lip 5-9 mm; anthers free; pistil included; stigmas 2; lateral veins 6-13 on each side of midrib; bracts linear.   (5)
       
5 (4) Leaf blade abaxially densely woolly on entire surface, 7.5-13 cm, base cuneate; lateral veins 8-13 on each side of midrib; peduncle 8.5-13 cm   1 Opithandra sinohenryi
+ Leaf blade abaxially appressed puberulent, woolly only along veins, 2.5-8 cm, base rounded to cordate; lateral veins 6-8 on each side of midrib; peduncle 2-6 cm   2 Opithandra burttii
       
6 (2) Leaf blade margin nearly entire to indistinctly repand; calyx segment apex caudate-acuminate, tip somewhat obtuse; corolla outside glabrous   6 Opithandra dinghushanensis
+ Leaf blade margin denticulate to serrulate to crenate or double crenate; calyx segment apex acute to obtuse; corolla outside sparsely puberulent to strigose at least adaxially.   (7)
       
7 (6) Leaf blade 3-19 cm, base cordate to broadly cuneate; lateral veins 7-9 on each side of midrib; peduncle 4.5-7.5 cm; corolla mouth 1.5-1.7 cm in diam.; stamens adnate to corolla near base, ca. 2 cm; pistil densely glandular puberulent to puberulent   5 Opithandra dalzielii
+ Leaf blade to 9.5 cm, base cuneate to broadly so; lateral veins 4-6 on each side of midrib; peduncle 5.5-14 cm; corolla mouth 5-9 mm in diam.; stamens adnate to corolla tube slightly below middle, 5-7.5 mm; pistil glabrous.   (8)
       
8 (7) Petiole 1.3-7.5 cm, with dense, white to brownish shorter appressed hairs and brown longer spreading ones; leaf blade 4-9.5 cm; corolla ca. 3.2 cm, mouth ca. 9 mm in diam.; staminodes 3   7 Opithandra obtusidentata
+ Petiole 0.3-1.7 cm, densely grayish strigose; leaf blade 1.5-3.5 cm; corolla 1.5-2 cm, mouth 5-6 mm in diam.; staminodes 2   8 Opithandra cinerea

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