1 |
Plants dioecious |
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(2) |
+ |
Florets bisexual |
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(3) |
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2 (1) |
Stems glabrous or cobwebby-felted; leaves divided or undivided but if undivided then base cuneate. |
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45 C. arvense |
+ |
Stems hairy; leaves undivided with base auriculate and semiamplexicaul. |
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46 C. shihianum |
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3 (1) |
Some of phyllaries expanded laterally into membranous wings or distally into a scarious appendage |
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(4) |
+ |
Phyllaries lacking wings and scarious appendage |
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(12) |
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4 (3) |
Leaves adaxially rough and spinulose |
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(5) |
+ |
Leaves adaxially smooth or covered with hair-tipped papillae and slightly rough but not spinulose |
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(6) |
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5 (4) |
Leaves abaxially densely felted; outer and middle phyllaries unwinged but inner phyllaries expanded into a scarious white or pink appendage. |
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37 C. leducii |
+ |
Leaves abaxially sparsely cobwebby; outer and middle phyllaries with a margin expanded into yellowish scarious wings. |
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44 C. chrysolepis |
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6 (4) |
Bracts surrounding capitula with pectinately arranged long pungent spines |
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(7) |
+ |
Bracts surrounding capitula (if present) without pectinately arranged long pungent spines |
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(9) |
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7 (6) |
Margin of middle phyllaries not laterally expanded into wings but inner phyllaries apically expanded. |
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41 C. argyracanthum |
+ |
Margin of middle phyllaries expanded above base into denticulate to lacerate scarious wings |
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(8) |
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8 (7) |
Outer phyllaries unwinged but usually with pectinately spinulose margin, middle phyllaries with irregularly lacerate scarious wings. |
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42 C. glabrifolium |
+ |
Outer and middle phyllaries with regularly denticulate-lacerate wings. |
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43 C. sieversii |
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9 (6) |
Leaves green, concolorous, and glabrous or with sparse hairs along veins |
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(10) |
+ |
Leaves abaxially grayish white and cobwebby or felted, adaxially green, glabrous, papillose, or with scattered crispate hairs |
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(11) |
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10 (9) |
Stems below capitulum felted. |
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33 C. tianmushanicum |
+ |
Stems with crispate hairs, below capitulum also ± cobwebby. |
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34 C. chinense |
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11 (9) |
Leaves undivided or rarely slightly lobed in proximal third. |
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35 C. lineare |
+ |
Leaves pinnately lobed to pinnatipartite at least in proximal half. |
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36 C. shansiense |
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12 (3) |
Outer and middle phyllaries villous or hirsute with multicellular hairs or bristles |
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(13) |
+ |
Involucre glabrous, sparsely cobwebby, or lanate, phyllaries never hirsute or villous |
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(14) |
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13 (12) |
Leaves concolorous, greenish, with sparse to dense bristles. |
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15 C. fuscotrichum |
+ |
Leaves discolorous, abaxially grayish white and densely felted, adaxially green and glabrous. |
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16 C. muliense |
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14 (12) |
Involucre densely lanate |
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(15) |
+ |
Involucre glabrous or cobwebby |
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(16) |
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15 (14) |
Leaves adaxially with scattered spinules or smooth, pinnately divided, segments elliptic, semiorbicular, or ovate. |
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7 C. eriophoroides |
+ |
Leaves adaxially rough with sparse to dense spinules, bipinnatipartite, segments triangular to triangular-lanceolate. |
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12 C. lidjiangense |
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16 (14) |
Leaves adaxially rough and spinulose |
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(17) |
+ |
Leaves adaxially smooth and not spinulose |
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(22) |
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17 (16) |
Stems winged. |
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14 C. vulgare |
+ |
Stems unwinged |
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(18) |
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18 (17) |
Leaf segments elliptic, semiorbicular, or ovate |
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(19) |
+ |
Leaf segments lanceolate to narrowly triangular |
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(20) |
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19 (18) |
Leaves discolorous, abaxially grayish white, adaxially green. |
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8 C. subulariforme |
+ |
Leaves concolorous, green. |
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9 C. verutum |
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20 (18) |
Leaves concolorous, green to yellowish green. |
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10 C. chlorolepis |
+ |
Leaves discolorous, abaxially grayish white, adaxially greenish |
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(21) |
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21 (20) |
Outer and middle phyllaries ending in a subulate spine 8-13 mm, outer phyllaries not much shorter than inner. |
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11 C. botryodes |
+ |
Outer phyllaries ending in a spinule or acumen to 2 mm, outer phyllaries much shorter than inner. |
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13 C. interpositum |
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22 (16) |
Corolla yellow or white |
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(23) |
+ |
Corolla red to purple |
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(24) |
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23 (22) |
Plants 80-150 cm tall; involucre 4-4.5 cm in diam.; leaves concolorous. |
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6 C. monocephalum |
+ |
Plants 45-50 cm tall; involucre ca. 2 cm in diam.; leaves discolorous. |
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40 C. alberti |
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24 (22) |
Plants stemless with all leaves basal |
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(25) |
+ |
Plants with leafy stems at least 20 cm tall |
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(26) |
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25 (24) |
Petiole unwinged or with discontinuous wing; outer and middle phyllaries narrowly triangular, pectinately fringed with spinules. |
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4 C. souliei |
+ |
Petiole with continuous wing; outer and middle phyllaries ovate to lanceolate, margin not spinulose. |
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23 C. esculentum |
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26 (24) |
Stems winged. |
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29 C. alatum |
+ |
Stems unwinged |
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(27) |
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27 (26) |
Outer phyllaries bractlike, dissimilar from and almost twice as long as middle and inner phyllaries. |
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1 C. bracteiferum |
+ |
Outer phyllaries not bractlike, similar to and not longer than middle ones |
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(28) |
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28 (27) |
Outer and middle phyllaries pectinately fringed with spinules |
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(29) |
+ |
Outer and middle phyllaries not fringed with spinules |
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(31) |
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29 (28) |
Leaves abaxially cobwebby or glabrous, adaxially glabrous. |
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5 C. fargesii |
+ |
Leaves not cobwebby but with crispate multicellular hairs |
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(30) |
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30 (29) |
Involucre ca. 4 cm in diam., sparsely cobwebby. |
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2 C. leo |
+ |
Involucre ca. 2 cm in diam., glabrous. |
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3 C. henryi |
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31 (28) |
Bracts surrounding capitula reduced to pectinately arranged long pungent spines |
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(32) |
+ |
Bracts surrounding capitula (if present) not reduced to pectinately arranged long pungent spines |
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(34) |
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32 (31) |
Leaves concolorous, green, glabrous. |
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38 C. semenowii |
+ |
Leaves discolorous, abaxially grayish green to whitish and cobwebby or felted, adaxially greenish and hairy |
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(33) |
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33 (32) |
Outer and middle phyllaries narrowed into a 0.1-0.5 cm spine. |
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24 C. periacanthaceum |
+ |
Outer and middle phyllaries tipped with a ca. 3 cm spine. |
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39 C. sairamense |
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34 (31) |
Phyllaries in ca. 10 rows; corolla tube more than twice as long as limb. |
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32 C. pendulum |
+ |
Phyllaries in 5-8 rows; corolla tube shorter or at most slightly longer than limb |
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(35) |
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35 (34) |
Leaves undivided, margins entire or toothed with spinules of at most 2 mm |
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(36) |
+ |
Leaves pinnately divided (or at least lobed), margin usually distinctly spiny |
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(41) |
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36 (35) |
Outer and middle phyllaries with an apical spine 3-7 mm. |
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30 C. vernonioides |
+ |
Phyllaries with an apical spinule of at most 1 mm |
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(37) |
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37 (36) |
Leaves discolorous, abaxially grayish to white and felted |
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(38) |
+ |
Leaves concolorous, abaxially green and glabrous or with multicellular hairs but not felted |
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(39) |
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38 (37) |
Lower cauline leaves petiolate, to 3 cm wide, narrowed at base. |
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19 C. vlassovianum |
+ |
All cauline leaves sessile, at least 4 cm wide, semiamplexicaul. |
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21 C. helenioides |
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39 (37) |
Involucre ovoid. |
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18 C. serratuloides |
+ |
Involucre campanulate |
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(40) |
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40 (39) |
Outer and middle phyllaries abaxially with a dark resinous gland. |
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17 C. viridifolium |
+ |
Phyllaries lacking a resinous gland. |
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20 C. fanjingshanense |
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41 (35) |
At least upper cauline leaves discolorous, abaxially gray to white and felted, adaxially ± green |
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(42) |
+ |
Leaves concolorous, abaxially green and glabrous or with sparse to dense multicellular hairs but not felted |
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(43) |
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42 (41) |
Lower cauline leaves petiolate, often concolorous; capitula corymbose. |
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25 C. maackii |
+ |
All cauline leaves sessile, discolorous; capitula racemose. |
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26 C. racemiforme |
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43 (41) |
Involucre 2-3 cm in diam |
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(44) |
+ |
Involucre 3-4.5 cm in diam |
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(45) |
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44 (43) |
Cauline leaves sessile; basal leaves present at anthesis. |
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22 C. japonicum |
+ |
Lower and middle cauline leaves petiolate; basal leaves withered at anthesis. |
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28 C. schantarense |
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45 (43) |
Corolla tube less than 1/2 as long as limb; outer and middle phyllaries abaxially with a dark resinous gland. |
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27 C. fangii |
+ |
Corolla tube slightly longer than limb; phyllaries lacking a resinous gland. |
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31 C. handelii |