| 1 |
Ovary with 1 ovule; fruit an achene. |
|
(2) |
| + |
Ovary with several or many ovules; fruit a follicle, rarely a capsule or berry. |
|
(17) |
| |
|
|
|
| 2 (1) |
Inflorescence with an involucre. |
|
(3) |
| + |
Inflorescence without an involucre. |
|
(6) |
| |
|
|
|
| 3 (2) |
Involucre contiguous with flower, calyxlike |
|
25 Hepatica |
| + |
Involucre separate from flower. |
|
(4) |
| |
|
|
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| 4 (3) |
Leaves pinnatisect, penninerved; pollen grains spinulose |
|
26 Anemoclema |
| + |
Leaves 1 to few times palmately or subpinnately divided, palmately nerved; pollen grains not spinulose. |
|
(5) |
| |
|
|
|
| 5 (4) |
Style strongly elongate and plumose in fruit |
|
27 Pulsatilla |
| + |
Style not as above |
|
24 Anemone |
| |
|
|
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| 6 (2) |
Petals absent. |
|
(7) |
| + |
Petals present. |
|
(10) |
| |
|
|
|
| 7 (6) |
Sepals valvate in bud; style usually strongly elongate and plumose in fruit; cauline leaves opposite, rarely alternate |
|
29 Clematis |
| + |
Sepals imbricate in bud; style not as above; cauline leaves (if present) alternate. |
|
(8) |
| |
|
|
|
| 8 (7) |
Achene (1--)3-ribbed on each side; leaves compound, rarely simple |
|
21 Thalictrum |
| + |
Achene not ribbed; leaves simple. |
|
(9) |
| |
|
|
|
| 9 (8) |
Basal leaves several, palmately lobed or undivided, veins reticulately anastomosing; staminodes absent |
|
28 Metanemone |
| + |
Basal leaf 1, palmatisect, veins dichotomously branched, not anastomosing; staminodes present |
|
31 Kingdonia |
| |
|
|
|
| 10 (6) |
Vines woody; cauline leaves opposite; style strongly elongate and plumose in fruit |
|
30 Naravelia |
| + |
Herbs perennial or annual; cauline leaves (if present) alternate; style not as above. |
|
(11) |
| |
|
|
|
| 11 (10) |
Petals without nectaries |
|
33 Adonis |
| + |
Petals with nectaries. |
|
(12) |
| |
|
|
|
| 12 (11) |
Aggregate fruit terete, achene with a robust, rostrate, persistent style at apex and 2 tubercles near base |
|
38 Ceratocephala |
| + |
Aggregate fruit not terete, achene with neither robust, rostrate, persistent style at apex nor tubercles near base. |
|
(13) |
| |
|
|
|
| 13 (12) |
Basal leaves several times pinnatisect; ovule pendulous from ovary locule apex |
|
32 Callianthemum |
| + |
Basal leaves not pinnatisect; ovule attached to ovary locule base. |
|
(14) |
| |
|
|
|
| 14 (13) |
Achene ribbed on each side. |
|
(15) |
| + |
Achene not ribbed. |
|
(16) |
| |
|
|
|
| 15 (14) |
Stolons absent; petals 13--15; achene 1-ribbed on each side |
|
36 Oxygraphis |
| + |
Stolons present; petals 5--10; achene 2- or 3-ribbed on each side |
|
37 Halerpestes |
| |
|
|
|
| 16 (14) |
Petals yellow; achene smooth, rarely spinulose or tuberculate |
|
34 Ranunculus |
| + |
Petals white, rarely yellow; achene transversely rugose on each side |
|
35 Batrachium |
| |
|
|
|
| 17 (1) |
Flowers zygomorphic; inflorescences usually racemose; pedicels 2-bracteolate. |
|
(18) |
| + |
Flower actinomorphic; inflorescences monochasial or, if racemose, pedicels ebracteolate. |
|
(20) |
| |
|
|
|
| 18 (17) |
Upper sepal spurless; petals clawed |
|
10 Aconitum |
| + |
Upper sepal spurred; petals clawless. |
|
(19) |
| |
|
|
|
| 19 (18) |
Staminodes 2, each differentiated into claw and limb; petals free; carpels 3--5(--7) |
|
11 Delphinium |
| + |
Staminodes absent; petals connate; carpel 1 |
|
12 Consolida |
| |
|
|
|
| 20 (17) |
Flowers many, borne in compound cymes, panicles, or racemes. |
|
(21) |
| + |
Flowers solitary, terminal, or few in monochasia. |
|
(24) |
| |
|
|
|
| 21 (20) |
Leaves simple, undivided; petals and staminodes absent |
|
4 Beesia |
| + |
Leaves 1 or more times ternately or subpinnately compound; petals or staminodes present. |
|
(22) |
| |
|
|
|
| 22 (21) |
Proximal cauline leaves sheathlike; sepals 8--11 mm; fruit slender stipitate |
|
5 Souliea |
| + |
Proximal cauline leaves not sheathlike; sepals less than 4.5 mm; fruit shortly stipitate or sessile. |
|
(23) |
| |
|
|
|
| 23 (22) |
Fruit a berry; basal leaves scalelike |
|
7 Actaea |
| + |
Fruit a follicle; basal leaves not scalelike |
|
6 Cimicifuga |
| |
|
|
|
| 24 (20) |
Petals absent. |
|
(25) |
| + |
Petals present. |
|
(27) |
| |
|
|
|
| 25 (24) |
Leaves 2-ternate |
|
14 Enemion |
| + |
Leaves simple. |
|
(26) |
| |
|
|
|
| 26 (25) |
Leaves entire |
|
1 Caltha |
| + |
Leaves 3-sect |
|
2 Calathodes |
| |
|
|
|
| 27 (24) |
Carpels connate at base |
|
22 Dichocarpum |
| + |
Carpels free. |
|
(28) |
| |
|
|
|
| 28 (27) |
Petals long clawed. |
|
(29) |
| + |
Petals very shortly clawed or clawless. |
|
(30) |
| |
|
|
|
| 29 (28) |
Leaves simple, peltate, entire or palmately lobed; carpels 5--8, free |
|
13 Asteropyrum |
| + |
Leaves usually pedately compound, rarely ternate, not peltate; carpels 2, connate at base. |
|
(28) |
| |
|
|
|
| 30 (28) |
Carpels slender stipitate |
|
23 Coptis |
| + |
Carpels sessile. |
|
(31) |
| |
|
|
|
| 31 (30) |
Staminodes present, membranous, linear-lanceolate. |
|
(32) |
| + |
Staminodes absent. |
|
(34) |
| |
|
|
|
| 32 (31) |
Sepals white; stamens 8--14; style 1/6--1/5 as long as ovary |
|
20 Semiaquilegia |
| + |
Sepals blue or purple; stamens numerous; style ca. 1/2 as long as to longer than ovary. |
|
(33) |
| |
|
|
|
| 33 (32) |
Leaves all basal, simple, 3-sect, or ternate; petals ca. 1/3 as long as sepals, not petaloid (instead specialized nectariferous organs); style ca. 2 × as long as ovary; follicles cubic-globose |
|
18 Urophysa |
| + |
Leaves both basal and cauline, 2 or more times ternate; petals subequaling sepals, petaloid; style ca. 1/2 as long as ovary; follicles narrowly cylindric |
|
19 Aquilegia |
| |
|
|
|
| 34 (31) |
Leaves simple. |
|
(35) |
| + |
Leaves compound. |
|
(37) |
| |
|
|
|
| 35 (34) |
Leaves all basal; inflorescence with an involucre |
|
9 Eranthis |
| + |
Leaves both basal and cauline; inflorescence without an involucre. |
|
(36) |
| |
|
|
|
| 36 (35) |
Leaves palmatipartite or palmatisect; petals linear |
|
3 Trollius |
| + |
Leaves pedatisect; petals tubular or cupular |
|
8 Helleborus |
| |
|
|
|
| 37 (34) |
Leaves all basal, persistent remnants of petioles forming a dense clump; flower solitary, terminal; petals clawless |
|
17 Paraquilegia |
| + |
Leaves both basal and cauline, petioles not forming a clump; flowers few, borne in a monochasium; petals very shortly clawed. |
|
(38) |
| |
|
|
|
| 38 (37) |
Herbs perennial; carpels 1--5 |
|
15 Isopyrum |
| + |
Herbs annual; carpels 6--20 |
|
16 Leptopyrum |