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FOC | Family List | FOC Vol. 20-21 | Asteraceae

145. Cotula Linnaeus, Sp. Pl. 2: 891. 1753.

山芫绥属 shan yuan sui shu

Authors: Zhu Shi, Christopher J. Humphries & Michael G. Gilbert

Lancisia Fabricius; Machlis Candolle.

Herbs, annual or perennial; indumentum absent or of basifixed hairs. Leaves alternate or sometimes opposite or rosulate, pinnatisect, lobed, or occasionally entire. Capitula solitary, pedunculate, heterogamous, radiate, rarely homogamous and discoid. Involucres hemispheric; phyllaries in 2 or 3 rows, unequal, scarious margin narrow. Receptacle flat to conical, epaleate. Marginal florets in 1 to several rows, female, fertile, generally pedicellate, corolla tube short or absent, lamina generally absent, white if present; disk florets yellow, bisexual, fertile, apex 4- or 5-lobed. Anther bases obtuse; apical appendage lanceolate. Style branches truncate or obtuse at apex or style undivided. Achenes dorsiventrally flattened, often laterally winged, wings aristately lengthened at apex of achene. Corona absent.

Fifty-five species: mostly in S Hemisphere, particularly S Africa and the Pacific islands (New Zealand), also in Australia and South America, a few species extending to E Africa and New Guinea; two species in China.

1 Achenes of marginal female florets obovoid; lateral wings broad and thick; ultimate segments of leaves shortly triangular or lanceolate.   1 C. anthemoides
+ Achenes of marginal female florets narrowly oblong; lateral wings very narrow; ultimate segments of leaves linear or linear-lanceolate.   2 C. hemisphaerica

Lower Taxa


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