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FOC | Family List | FOC Vol. 13 | Clusiaceae

4. Cratoxylum Blume, Verh. Batav. Genootsch. Kunsten. 9: 172, 174. 1823.

黄牛木属 huang niu mu shu

Authors: Xi-wen Li, Jie Li & Peter Stevens

Hornschuchia Blume, Catalogus, 15. 1823, not Nees (1821).

Trees or shrubs, deciduous to evergreen. Twigs with nodes sometimes flattened, mostly with interpetiolar lines; apical bud abortive or not; buds with scales or not. Leaves opposite, sessile to petiolate, entire, abaxially often pruinose to waxy, often finely translucent-punctate between veins, secondary veins ± widely spaced, tertiary veins reticulate. Inflorescence cymose, terminal and/or axillary; bracteoles caducous, minute. Flowers bisexual, homostylous or heterostylous, pedicellate. Sepals 5, ± unequal, leathery, usually accrescent. Petals 5, white to deep crimson or pink, sometimes tinged orange and green, often with punctiform or linear dark resinous glands, sometimes with ± adnate adaxial near-basal scalelike appendage. Stamen fascicles apparently 3 (2 + 2 + 1), with filaments ca. 2/3 united; anthers subpersistent, dorsifixed, sometimes with ± prominent resinous connective gland. Fasciclodes 3, alternating with fascicles, scalelike, sometimes minute. Ovary 3-loculed, with [3-]5-16[-18] erect or ascending ovules on lower half of each placenta; styles 3, free, often divergent; stigmas punctiform, truncate or somewhat thickened, slightly papillose. Capsule ± woody, ellipsoid to narrowly cylindric, dehiscing loculicidally, with persistent columella. Seeds obovoid to cylindric, with unilateral wing 2.5-3 × as long [or wing all around and smaller]; embryo cylindric.

About six species: Cambodia, S and SW China, India, Indonesia, Laos, Malaysia, Myanmar, Philippines, Thailand, Vietnam, all S of ca. 24° N; two species in China.

1 Leaves with secondary veins ascending, not or irregularly joining; inflorescences terminal and axillary; flowers homostylous; stamen fascicles short and broad; fasciclodes recurved-cucullate; twigs and leaves glabrous; petals without appendage.   1 C. cochinchinense
+ Leaves with secondary veins spreading, regularly joining toward margin; inflorescences axillary, often on old growth; flowers heterostylous; stamen fascicles long and slender; fasciclodes never recurved; twigs and leaves sometimes villous; petals with minute, basal appendage.   2 C. formosum

Lower Taxa


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