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FOC | Family List | FOC Vol. 2-3 | Pteridaceae

2. Cryptogramma R. Brown in Franklin, Narr. Journey Polar Sea. 767. 1823.

珠蕨属 zhu jue shu

Authors: Zhang Gangmin, Edward R. Alverson & Jordan S. Metzgar

Cryptogramma raddeana

Credit: Harvard University Herbaria

Plants small, usually on rocks. Rhizomes short, erect or ascending with dictyostele, or occasionally long creeping with siphonostele; scales colorless or brownish, thin. Fronds strongly dimorphic, clustered or rarely scattered, sterile fronds shorter than fertile ones; stipe dark brown proximally, light brown to straw-colored distally, grooved adaxially, scaly. Sterile fronds: lamina broadly ovate or oblong, 2-4-pinnate, membranous to leathery, glabrous. Fertile fronds: lamina 2- or 3-pinnate. Ultimate segments of sterile lamina ovate, spatulate, elliptic, or fan-shaped; fertile segments linear or elongate oblong. Veins free, pinnately branched, simple or forked distally; hydathodes often sunken below surface on dried specimens. False indusia broad, clearly marginal, continuous, making a single fertile segment podlike. Sori borne at vein tips, orbicular or elliptic, confluent when mature. Spores yellow, tetrahedral, trilete, verrucose. x = 30.

About ten species: temperate and subtropical alpine regions, Asia, Europe, North and South America; three species in China.

Plants of Cryptogramma resemble those of Onychium but are smaller, strongly dimorphic, and have fertile fronds markedly taller than sterile fronds. There is not a commisural vein connecting veinlet tips, so the sori are borne at veinlet tips and become confluent only when mature.

1 Rhizomes slender and creeping; fronds sparse and scattered; sterile lamina 1- or 2-pinnate, herbaceous to membranous, thin; hydathodes absent.   1 C. stelleri
+ Rhizomes stout and erect; fronds densely tufted; sterile lamina 3- or 4-pinnate, papery or somewhat leathery, obscure; hydathodes present and conspicuous   (2)
2 (1) Sterile lamina 3-pinnate-pinnatifid, ultimate segments triangular, with an acute apex.   2 C. brunoniana
+ Sterile lamina 4-pinnate-pinnatifid, ultimate segments linear, with a rounded apex.   3 C. raddeana

Lower Taxa


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