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17. Dactylicapnos Wallich, Tent. Fl. Napal. 2: 51. 1826.
紫金龙属 zi jin long shu
Authors: Mingli Zhang & Magnus Lidén
Herbs, scandent, perennial or annual, glabrous. Stems thin, 1-8 m, forming a sympodium of several shoots. Leaf blade ternately or pinnately compound; leaflets once to 3 × compound; apex of leaf transformed into a branched tendril (except in basalmost leaves). Inflorescences terminal (leaf-opposed), corymbose, pendent, 2-14-flowered; bracts linear, entire to fimbriate-dentate. Flowers with 2 planes of symmetry, cordate to oblong, 12-26 × 4-10 mm. Petals pale yellow to orange (often rubescent); apices of outer petals slightly bent outward, base pouchlike. Stamens 2 (i.e., filaments of each triplet completely fused); each stamen with a basal nectary extending into pouch of corresponding outer petal. Style persistent; stigma almost square, upper corners each with 1 small stigmatic papilla, basal corners each with 1 large papilla (small in Dactylicapnos burmanica). Capsule dehiscent with 2 valves with persistent replum (in D. scandens indehiscent and berrylike). Seeds several to many, carunculate.
Twelve species: Himalayas to W China; ten species (three endemic) in China.
Dactylicapnos ventii (Khánh) Lidén (Nordic J. Bot. 25: 35. 2008) is similar to D. grandifoliolata but has fruit lanceolate with thin walls and seeds evenly colliculate and slightly flattened. It is known from India (Sikkim) and might occur in Yadong, S Xizang.
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1 |
Bracts and sepal margins fimbriate-dentate; plant a squashy annual; stems weak, sharply angular or winged (D. sect. Minicalcara (Khanh) Lidén) |
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(2) |
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Bracts entire, sepals subentire; leaves (tendril excepted) with 2 primary leaflets (rarely 4, but then opposite); plant a perennial; stems terete to sulcate, but not winged (D. sect. Dactylicapnos) |
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(5) |
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2 (1) |
Flowers sharply keeled, 8-13 mm wide; stems conspicuously winged-ridged. |
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7 D. roylei |
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Flowers not keeled, 4-6 mm wide; stems with narrow ridges |
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(3) |
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3 (2) |
Capsule constricted between seeds; seeds shiny, radially reticulate-striate; first flowers 1-2 m up stem. |
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8 D. torulosa |
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Capsule smooth; seeds smooth or densely echinulate; first flowers usually 0.1-1 m up stem |
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(4) |
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4 (3) |
Seeds echinulate; sinus between basal pouches to 1 mm deep. |
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9 D. lichiangensis |
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Seeds smooth; sinus between basal pouches ca. 2 mm deep. |
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10 D. leiosperma |
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5 (1) |
Primary leaflets 2, alternate; nectary even or tapering toward apex |
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(6) |
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Primary leaflets 2 or 4, opposite; nectary much thicker toward apex |
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(7) |
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6 (5) |
Capsule very fleshy, not dehiscing by regular valves; nectary sigmoid, even or apex shortly acute. |
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1 D. scandens |
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Capsule membranous with thin marginal wing, dehiscing with valves; nectary sharply geniculate, apex long and thin. |
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2 D. macrocapnos |
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7 (5) |
Primary leaflets bi- to triternate, each with 8-16 ultimate leaflets; seeds with small elaiosomes; fruit 3-6 × as long as wide |
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(8) |
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Primary leaflets each with 2-8 ultimate leaflets; seeds with large deeply dissected elaiosomes that keep them together in a clump after valves open; fruit 1.7-2.5 × as long as wide (longer in D. ventii) |
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(9) |
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8 (7) |
Most leaves with 2 pairs of lateral leaflets; stems 2-4 mm in diam.; nectary not T-shaped; stigma broader than long with large basal papillae; crests of inner petals overtopping apex by 2 mm. |
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3 D. schneideri |
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Leaves with 1 pair of lateral leaflets; stems 1-2 mm in diam.; nectary T- or Y-shaped; stigma longer than broad with small basal papillae; crests of inner petals overtopping apex by 4 mm. |
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4 D. burmanica |
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9 (7) |
Primary leaflets each with 4-8 obtuse to subacute ultimate leaflets; fruit with 15-30 seeds. |
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5 D. gaoligongshanensis |
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Primary leaflets each with 3-5 acute to subacute ultimate leaflets; fruit with 30-50 seeds |
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(10) |
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10 (10) |
Fruit thin-walled, lanceolate; seeds colliculate. ..... D. ventii (see note above) |
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(10) |
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Fruit thick-walled, broadly elliptic; seeds echinulate. |
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6 D. grandifoliolata |
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Lower Taxa
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