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FOC | Family List | FOC Vol. 19 | Dipsacaceae

4. Dipsacus Linnaeus, Sp. Pl. 1: 97. 1753.

川续断属 chuan xu duan shu

Herbs, biennial or perennial. Stems erect, angulate and sulcate, usually echinulate or hispid along ridges. Basal leaves long petiolate, entire, 3-cleft, or pinnatipartite, usually dentate or lobed. Cauline leaves opposite, sessile or petiolate, usually 3-5-lobed, sometimes pinnatipartite or entire, both surfaces usually hispid, but glabrous or papillate-echinulate in some species. Capitula terminal, oblong, globose, or ovoid-globose; involucral bracts erect or spreading, leaflike, 1- or 2-layered, rostrate; involucels 4-8-ribbed, adnate to ovary; flowers bisexual. Calyx actinomorphic, shallowly disciform, 4-lobed, white pubescent. Corolla tubular, 4-lobed; lobes unequal in size. Stamens 4, inserted at corolla tube, alternate with corolla lobes. Gynoecium of 2 carpels; ovary inferior, enveloped by involucel; ovule solitary, pendulous, anatropous; style filiform; stigma oblique or lateral. Achenes surmounted with persistent calyx, enveloped by leathery involucel. Seed coat membranous; embryo imbedded in endosperm.

About 20 species: N Africa, Asia, Europe; seven species (two endemic) in China.

1 Stems glabrous; bract rostra 1.5-3 mm.   1 D. azureus
+ Stems ± spiny; bract rostra 3-11 mm   (2)
2 (1) Heads oblong-spherical, 2.5-7 × 2-4.9 cm; bract rostra 5-11 mm; corolla tube 3.5-5 mm   (3)
+ Heads spherical, 1.5-3 × 1.5-3.2 cm; bract rostra 3-5 mm; corolla tube 2-2.5 mm   (5)
3 (2) Corolla black-purple, tube ca. 3.5 mm; bract rostra ca. 5 mm.   4 D. atratus
+ Corolla white or yellowish, tube 3.5-5 mm; bract rostra 5-11 mm   (4)
4 (3) Segments of cauline leaves 1-3-paired; heads 2-3.5 cm in diam.; leaves hirsute, especially along veins, abaxially   2 D. inermis
+ Segments of cauline leaves more than 3-paired; heads usually 3.5-4.9 cm in diam.; leaves spiny abaxially.   3 D. chinensis
5 (2) Corolla black-purple.   6 D. atropurpureus
+ Corolla white, yellow, or pink   (6)
6 (5) Corolla white or yellow; plant perennial; roots fleshy and fusiform.   5 D. asper
+ Corolla pink; plant biennial; roots ± lignified, not fleshy.   7 D. japonicus

Lower Taxa


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