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FOC | Family List | FOC Vol. 22 | Poaceae

131. Eragrostis Wolf, Gen. Pl. 23. 1776.

画眉草属 hua mei cao shu

Authors: Shou-liang Chen & Paul M. Peterson

Annual or perennial, often glandular particularly on the leaf sheaths and inflorescence. Leaf blades mostly flat, sometimes rolled, rarely pungent; ligule a line of hairs or sometimes membranous. Inflorescence an open, contracted, spiciform or glomerate panicle, very rarely of racemes on a central axis. Spikelets 2- to many-flowered, laterally compressed, orbicular to vermiform, variously disarticulating. Glumes unequal, deciduous or persistent, 1(–3)-veined. Lemmas membranous to coriaceous, keeled or rounded, glabrous to asperulous or rarely hairy, 3-veined or the veins sometimes very faint and occasionally suppressed, apex entire, obtuse to acuminate, rarely mucronate. Palea keels sometimes winged or ciliate. Stamens 2 or 3. Fruit mostly globose, ellipsoid, or rectangular-prismatic, usually a caryopsis but sometimes the pericarp free.

About 350 species: tropics and subtropics throughout the world; 32 species (11 endemic, one introduced) in China.


1 Florets disarticulating from above downward, falling together with the rachilla joints   (2)
+ Florets disarticulating from below upward, usually leaving the rachilla entire   (7)
       
2 (1) Panicle contracted, spikelike or cylindrical   (3)
+ Panicle usually open   (4)
       
3 (2) Perennial; lemmas 1.8–2.5 mm, ciliolate along the margins below.   27 E. ciliata
+ Annual; lemmas 0.8–1.3 mm, glabrous or scabrous along the margins.   28 E. ciliaris
       
4 (2) Perennial.   29 E. collina
+ Annual   (5)
       
5 (4) Palea-keels long ciliate; branchlets and pedicels glandular.   30 E. tenella
+ Palea-keels glabrous to ciliolate; branchlets and pedicels eglandular   (6)
       
6 (5) Culms 120–150 cm tall, ca. 5 mm in diam.; panicle branches solitary or in pairs; spikelets yellowish green   31 E. alta
+ Culms 30–100 cm tall, 1.5–2.5 mm in diam., panicle branches clustered or verticillate; spikelets purplish at maturity   32 E. japonica
       
7 (1) Annual   (8)
+ Perennial   (17)
       
8 (7) Palea falling together with its lemma at maturity   (9)
+ Palea persistent or tardily falling   (11)
       
9 (8) Spikelets oblong, 5–10 × 2–4 mm, 10–20-flowered.   25 E. unioloides
+ Spikelets filiform, 11–25 × 1–2.5 mm, 10–40-flowered   (10)
       
10 (9) Upper glumes ca. 1 mm; lower lemma ca. 1.5 mm; palea ca. 1 mm; anthers ca. 0.2 mm.   20 E. multicaulis
+ Upper glumes 1.3–2.3 mm; lower lemma 1.8–2.2 mm; palea ca. 1.6–1.8 mm; anthers 0.7–0.9 mm.   7 E. atrovirens
       
11 (8) Plants glandular on the culms, leaf sheaths, and panicle   (12)
+ Plants eglandular   (14)
       
12 (11) Spikelets 2–3 mm broad, lower lemma 2.2–2.8 mm.   22 E. cilianensis
+ Spikelets 1.5–2.5 mm broad, lower lemma 1.5–2 mm   (13)
       
13 (12) Palea subequal to its lemma; midrib of lemma glandular.   23 E. minor
+ Palea shorter than its lemma; midrib of lemma eglandular.   24 E. suaveolens
       
14 (11) Lower glume 1-veined, 1–2 mm, the upper ca. 2 mm   (15)
+ Lower glume without a vein, less than 1 mm, the upper less than 1.4 mm   (16)
       
15 (14) Spikelets 5–20 × 2–2.5 mm, 8–40 florets.   16 E. cumingii
+ Spikelets 3–5 × ca. 2 mm, 3–10 florets.   21 E. autumnalis
       
16 (14) Summit of sheaths pilose; panicles 3.5–14 cm wide, axils pilose; pedicels as long or longer than the spikelets.   19 E. pilosa
+ Summit of sheaths glabrous; panicles 1.5–3 cm wide, axils glabrous; pedicels usually shorter than the spikelets.   20 E. multicaulis
       
17 (7) Lemma falling together with the palea at maturity   (18)
+ Palea persistent or tardily falling at maturity   (19)
       
18 (17) Spikelets 2–4 mm wide; stamens 2, anthers 0.2–0.5 mm; leaf blades sublanceolate, 3–6 mm broad.   25 E. unioloides
+ Spikelets 1.5–2.5 mm wide; stamens 3, anthers 0.7–0.9 mm; leaf blades linear, 2–4 mm broad.   7 E. atrovirens
       
19 (17) Panicle contracted and spikelike, less than 3(–5) cm wide   (20)
+ Panicle open, usually more than 3 cm wide   (22)
       
20 (19) Spikelets 1–2 mm broad; lower panicle branches (1.5–)3–8 cm.   4 E. nutans
+ Spikelets 2–3 mm broad; lower panicle branches 0.5–2.5 cm   (21)
       
21 (20) Palea apex acute, the keels ciliate but not winged; panicle 2–8 cm, pilose in axils.   5 E. cylindrica
+ Palea apex toothed, the keels winged, ciliolate along the wings; panicle 10–15 cm, glabrous in axils.   6 E. nevinii
       
22 (19) Branchlets and pedicels distinctly or obscurely glandular   (23)
+ Branchlets and pedicels eglandular   (24)
       
23 (22) Caryopsis rectangular-prismatic with a shallow adaxial groove, laterally compressed, 0.7–1.5 mm.   15 E. ferruginea
+ Caryopsis obovoid to ellipsoid, terete, without a groove, 0.7–0.9 mm.   13 E. perennans
       
24 (22) Branches densely spiculate to base   (25)
+ Branches naked at base   (26)
       
25 (24) Stamens 3, anthers 0.3–1.3 mm; caryopsis dark brown.   1 E. brownii
+ Stamens 2, anthers 0.2–0.3 mm; caryopsis cinnamon.   2 E. elongata
       
26 (24) Leaf blades long pilose on both surfaces   (27)
+ Leaf blades glabrous, pubescent or pilose in part   (28)
       
27 (26) Spikelets 7–14-flowered; caryopsis furrowed on one side, bluntly triangular in section.   10 E. pilosissima
+ Spikelets usually 7-flowered; caryopsis compressed, elliptical in section.   11 E. pilosiuscula
       
28 (26) Spikelets livid green, black-green, purplish black, or plumbeous; leaf sheaths compressed at the base   (29)
+ Spikelets purplish, yellowish, or greenish; leaf sheaths not compressed at the base   (34)
       
29 (28) Culms 80–120 cm tall; leaf blades flat or involute, up to 40 cm; panicle up to 40 cm   (30)
+ Culms 30–80 cm tall; leaf blades flat or involute, 5–25 cm, panicle up to 23 cm   (31)
       
30 (29) Caryopsis ellipsoid to obovoid, smooth and mostly translucent, dorsally compressed, adaxial surface sometimes with a shallow, broad groove, light brown.   9 E. curvula
+ Caryopsis rectangular-prismatic, faintly striate, laterally compressed, with a shallow, narrow groove, reddish brown   15 E. ferruginea
       
31 (29) Plants with scaly buds at the base; spikelets 4–13 mm, 6–24-flowered   (32)
+ Plants without scaly buds at the base; 3–12-flowered   (33)
       
32 (31) Lower glume ca. 1 mm; upper glume ca. 1.3 mm, 1–3-veined; caryopsis 0.8–1 mm.   16 E. cumingii
+ Lower glume ca. 1.2 mm; upper glume ca. 1.8 mm, 1-nerved; caryopsis 0.5–0.6 mm.   17 E. duricaulis
       
33 (31) Spikelets 2–2.5 mm wide, 5–10 mm; lemmas 2–2.2 mm; caryopsis elliptical, terete   14 E. nigra
+ Spikelets 1–1.5 mm wide, 3–6 mm; lemmas 2.4–3 mm; caryopsis rectangular-prismatic, laterally compressed.   15 E. ferruginea
       
34 (28) Palea tardily deciduous at maturity; spikelets 7–15 mm, 10–44-flowered.   18 E. hainanensis
+ Palea persistent; spikelets 0.5–20 mm, 5–60-flowered   (35)
       
35 (34) Axils of panicle and branches glabrous   (36)
+ Axils of panicle and branches pilose   (37)
       
36 (35) Spikelets 5–10 × ca. 2.5 mm, 5–15-flowered; anthers ca. 0.5 mm.   8 E. fauriei
+ Spikelets 5–25 × ca. 3 mm, 6–60-flowered; anthers ca. 0.3 mm.   12 E. perlaxa
       
37 (35) Culms 50–110 cm tall; panicle 20–35 cm; anthers ca. 1 mm.   13 E. perennans
+ Culms 20–60 cm tall; panicle 3–12 cm; anthers 0.2–0.4 mm   (38)
       
38 (37) Leaf blades pilose on adaxial surface; lemmas with reddish or yellowish lateral veins.   3 E. rufinerva
+ Leaf blades glabrous; lemmas with inconspicuous lateral veins.   26 E. montana

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