1 |
Plicae reduced to a small tooth or auricle on side of lobe, less than 1 mm. |
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(2) |
+ |
Plicae usually wider and/or longer than 1 mm. |
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(3) |
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2 (1) |
Corolla lobes as long as or longer than tube |
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I Otophora |
+ |
Corolla lobes shorter than tube (in G. prainii) |
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XI Sect. Chondrophyllae |
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3 (1) |
Corolla throat with a ring of multicellular hairs (not to be confused with fringed plicae between corolla lobes) |
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XII Sect. Fimbricorona |
+ |
Corolla throat naked. |
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(4) |
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4 (3) |
Style almost as long as or longer than ovary; calyx winged or sharply angled or if terete then stems, leaves, and calyces densely covered with purple and white pilosity. |
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(5) |
+ |
Style usually much shorter than ovary (about as long as ovary in some species of sects. Isomeria and Microspermae); calyx not winged, usually terete. |
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(6) |
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5 (4) |
Plants 5-12 cm tall, densely branched from base and toward stem apex; stem leaves ± crowded toward apex, leaf blades lanceolate; stamens straight; seed coat alveolate |
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VIII Microsperma |
+ |
Plants (5-)10-50 cm tall, not densely branched; stem leaves widely separated, leaf blades ovate-lanceolate, ovate, or cordate; stamens decurved; seed coat minutely to coarsely reticulate |
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IX Stenogyne |
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6 (4) |
Stigma lobes connate or contiguous, expanded into a discoid or funnelform structure; calyx completely enveloped by upper pair of broad obovate leaves |
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VI Phyllocalyx |
+ |
Stigma lobes free, recurved, linear to oblong; calyx usually not concealed by upper leaves or leaves narrower. |
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(7) |
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7 (6) |
Stamens unequal and plants perennial; flowers 4merous or if 5merous then corollas 3-4.5 cm … |
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III Kudoa |
+ |
Stamens equal or if unequal then plants annuals with solitary flowers (flowers of G. picta and G. yunnanensis in clusters) and corollas 0.6-2.5 mm; flowers 5-merous. |
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(8) |
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8 (7) |
Perennials; basal leaves persistent at flowering time, blades (excluding petiole) mostly over 3 cm and much larger than stem leaves or vegetative rosettes present along with flowering stems and their leaves distinctly larger than those of flowering stems. |
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(9) |
+ |
Annuals or if perennials then without conspicuously larger basal or rosette leaves (largest leaves at flowering time usually well above base of stem), and/or with all basal leaves less than 1.5 cm. |
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(13) |
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9 (8) |
Flowering stems arising singly from slender stolons, often creeping and rooting at lower nodes, without a well-developed caudex or stout rhizome and without a stout persistent taproot; larger leaves generally distinctly pseudopetiolate, with blades less than 1.8 cm, spatulate, elliptic, ovate-elliptic, obovate, or suborbicular, base obtuse, rounded, or truncate, only midvein prominent; flowers solitary or to 8 in terminal clusters, sessile |
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VII Sect. Isomeria |
+ |
Flowering stems arising from a ± stout erect caudex, often with a well-developed ± stout taproot and/or in a dense tuft (sometimes also stoloniferous); leaves and inflorescences diverse but above combination of characters generally not present. |
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(10) |
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10 (9) |
Remains of old leaf bases fibrous and conspicuous on caudex; seed coat reticulate, usually minutely so (also winged at 1 end in G. waltonii) |
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II Cruciata |
+ |
Remains of old leaf bases at base of current season's growth absent or membranous, not distinctly fibrous or if somewhat fibrous then with stem leaves numerous, closely spaced, less than 5 mm wide, and with flowers solitary; seed coat with membranous lamellae forming spongy complex hexagonal pits. |
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(11) |
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11 (10) |
Stems several to many, ± tufted, strongly decumbent; stem leaves opposite or whorled, numerous, closely spaced (9 or more pairs or whorls per stem and/or upper leaves exceeding internodes) |
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III Kudoa |
+ |
Stems 1 to several, decumbent or erect; stem leaves opposite, usually fewer than 9 pairs per stem, closely spaced only basally and on short stems. |
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(12) |
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12 (11) |
Sepals 3-9 mm, ± erect, 2 conspicuously larger than other three; base of current season's growth without blackish membranous remains of old leaf bases |
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III Kudoa |
+ |
Sepals smaller, often recurved or if over 3 mm then sepals subequal and/or base of current season's growth covered with blackish membranous remains of old leaf bases |
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IV Sect. Frigida |
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13 (8) |
Perennials, without a stout caudex; flowers solitary. |
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(14) |
+ |
Annuals or if perennial then with a stout caudex and clustered flowers; flowers solitary or clustered. |
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(15) |
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14 (13) |
Corolla over 2.5 cm, often much larger; capsule wingless |
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III Kudoa |
+ |
Corolla usually to 1.5 cm (2.3-3 cm in G. qiujiangensis and G. jamesii); capsule winged along sutures toward apex |
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XI Sect. Chondrophyllae |
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15 (13) |
Perennials; stems usually over 20 cm, from a stout caudex; larger stem leaf blades usually over 3.5 cm; flowers clustered. |
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(16) |
+ |
Annuals; stems usually shorter than 20 cm, caudex none; all stem leaf blades usually less than 3 cm; flowers solitary or clustered. |
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(17) |
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16 (15) |
Calyx lobes subequal or irregularly unequal, linear to narrowly triangular, base not or scarcely narrowed; calyx tube often split along 1 side (except in G. manshurica); lowermost stem leaves scalelike; seed coat coarsely reticulate, winged at ends |
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V Pneumonanthe |
+ |
Calyx lobes dimorphic, 2 larger than other 3, lanceolate to spatulate, base distinctly narrowed; calyx tube entire; lowermost stem leaves larger or scalelike; seed coat alveolate, not winged |
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III Kudoa |
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17 (15) |
Seed coat alveolate, with hexagonal pits; flowers solitary or clustered in axillary or terminal inflorescences, if clustered then with corollas over 1.5 cm and/or with calyx lobes each with distinct blade and petiole |
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VIII Microsperma |
+ |
Seed coat finely reticulate, without hexagonal pits; flowers usually solitary, if ± clustered in terminal inflorescences then with corolla 0.7-1.1 cm and calyx lobes without petiole. |
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(18) |
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18 (17) |
Capsule conspicuously winged along sutures at apex |
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XI Sect. Chondrophyllae |
+ |
Capsule not winged |
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X Sect. Dolichocarpa |
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