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161. Hippolytia Poljakov, Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 18. 288. 1957.

女蒿属 nü hao shu

Authors: Zhu Shi, Christopher J. Humphries & Michael G. Gilbert

Herbs, perennial, sometimes stemless, small subshrubs, or cushion plants; indumentum of medifixed hairs. Leaves alternate, pinnatifid or 3-lobed. Synflorescences cymose, dense or lax, forming panicles, clusters, or heads. Capitula few to many, homogamous, discoid. Involucre campanulate or conical; phyllaries in 3 or 4 rows, herbaceous or rigidly herbaceous, scarious margins white or brown. Receptacle flat to convex, epaleate. Florets all bisexual; corolla tubular, apex 5-lobed. Anther bases obtuse, apical appendage ovate-lanceolate. Style branches linear, apex truncate. Achenes subterete, 4-7-ribbed, with a ± distinct apical rim. Corona absent.

Nineteen species: C Asia, China, Himalaya, Mongolia; 11 species (six endemic) in China.

Hippolytia dolichophylla (Kitamura) K. Bremer & Humphries (Bull. Nat. Hist. Mus. London, Bot. 23: 106. 1993; Chrysanthemum dolichophyllum Kitamura, Acta Phytotax. Geobot. 23: 73. 1968, based on Tanacetum longifolium Wallich ex Candolle, Prodr. 6: 130. 1838, not Thunberg (1800); C. longifolium K. H. Rechinger (1955), not Sessé & Mociño (1894); T. dolichophyllum (Kitamura) Kitamura; T. himachalense Aswal & Mehrotra, nom. illeg. superfl.) was recorded from Sichuan in Annot. Checkl. Fl. Pl. Nepal (69. 2000, as T. dolichophyllum), disjunct from the main area of distribution in the Himalaya between Kashmir and Nepal (see; accessed 8 Apr 2011). No specimen has been seen by the present authors. This species would key out as H. herderi, but it differs from that species and all others in Hippolytia by having leaves 12-25 cm.

1 Rosette or cushion plants with concealed stems, to 7 cm tall, usually less; capitula in very dense sessile clusters   (2)
+ Plants with obvious stems, 7-50 cm tall; capitula in pedunculate panicles   (5)
2 (1) Rosette plants with a single central cluster of capitula   (3)
+ Cushion plants with several distinct clusters of capitula   (4)
3 (2) Leaves 2-pinnatisect.   8 H. glomerata
+ Leaves 3-7-pinnatisect.   9 H. syncalathiformis
4 (2) Leaves spatulate, apex 3-6-lobed.   10 H. gossypina
+ Leaves orbicular or flabelliform, biternate-palmately divided.   11 H. kennedyi
5 (1) Capitula fascicled-corymbose at apices of branches or stem; involucres campanulate, glossy, yellowish or straw-colored, phyllaries in 4 or 5 rows, scarious margins pale brownish or white   (6)
+ Capitula corymbose or glomerulate; involucres cuneate or cuneate-campanulate, matt, phyllaries in 3 or 4 rows, scarious margins dark brown   (8)
6 (5) Perennial herbs, often with a solitary stem.   3 H. delavayi
+ Subshrubs   (7)
7 (6) Leaves 2-pinnatisect, abaxially green or deep green, glabrous; florets ca. 3.2 mm.   2 H. desmantha
+ Leaves pinnatipartite, abaxially gray-white, densely appressed pubescent; florets ca. 2 mm.   1 H. kaschgarica
8 (5) Leaves 3-pinnatisect; capitula many.   4 H. yunnanensis
+ Leaves 2-pinnatisect; capitula 4-12   (9)
9 (8) Plants to 30 cm tall; leaves narrowly elliptic or ovate, 2-4 cm, ultimate segments narrowly elliptic.   5 H. herderi
+ Plants to 12 cm tall; leaves ovate, ultimate segments linear   (10)
10 (9) Stem leaves sessile.   6 H. senecionis
+ Stem leaves petiolate, petiole 5-10 mm.   7 H. tomentosa

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