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FOC | Family List | FOC Vol. 7 | Papaveraceae

13. Hypecoum Linnaeus, Sp. Pl. 1: 124. 1753.

角茴香属 jiao hui xiang shu

Authors: Mingli Zhang & Magnus Lidén

Hypecoum leptocarpum

Credit: Harvard University Herbaria

Chiazospermum Bernhardi.

Herbs, annual, low, usually glaucous, glabrous, taprooted, rosulate. Leaves in a basal rosette, lanceolate, oblanceolate, to narrowly obovate in outline, imparipinnate with deeply divided segments. Flowers in long pedunculate dichasia from rosette, bisymmetric. Petals 4, yellow, white, or bluish pink; outer petals flat, shallowly 3-lobed to entire; inner petals deeply 3-lobed; lateral lobes oblong, entire; median lobe with a narrow base and a rounded to oblong limb with a fimbriate or ± denticulate margin, specialized for secondary pollen presentation, initially pressed to stamens/gynoecium. Stamens 4, each with 2 thecae, but median ones with 2 vascular bundles, lateral ones with 1 vascular bundle; filaments flat, ± translucent, with lateral basal nectaries; anthers oblong, extrorse. Style with 2 narrow commissural stigmatic branches. Fruit linear, with many seeds in 1 row, dehiscing with 2 valves or breaking up into 1-seeded units. Seeds flattish, covered with oxalate crystals. Pollen with 2 long colpi, sometimes fused into a synaperture. Chromosome base number = 8.

Eighteen species: Mediterranean region to C Asia and China; four species (one endemic) in China.

See the paper by Dahl, "Infrageneric division of the genus Hypecoum L. (Papaveraceae)" (Nordic J. Bot. 10: 129-140. 1992).

1 Ultimate leaf lobes linear to setaceous; petals yellow   (2)
+ Ultimate leaf lobes obovate-lanceolate; outer petals white, pinkish, or bluish (H. sect. Leptocarpa Å. E. Dahl)   (3)
2 (1) Fruit dehiscent with 2 valves, ± erect; margin of central lobe of inner petals not fimbriate; dry plant not smelling of coumarin; outer petals > 10 mm wide (H. sect. Chiazospermum (Bernhardi) Å. E. Dahl).   1 H. erectum
+ Fruit nutant, breaking up into 1-seeded indehiscent units; margin of central lobe of inner petals fimbriate; dry plant smelling of coumarin; outer petals < 5 mm wide (H. sect. Pendula Å. E. Dahl).   4 H. parviflorum
3 (1) Outer petals 6-9 × 4-6 mm; anthers 1-1.5 mm; inner petals pale.   2 H. leptocarpum
+ Outer petals 11-12 × 9-10 mm; anthers 3-3.5 mm; inner petals ?orange.   3 H. zhukanum

Lower Taxa


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