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FOC | Family List | FOC Vol. 19 | Rubiaceae

44. Leptodermis Wallich in Roxburgh, Fl. Ind. 2: 191. 1824.

野丁香属 ye ding xiang shu

Authors: Tao Chen

Shrubs, sometimes low or reduced, unarmed, usually with short shoots. Raphides present. Leaves opposite, apparently without domatia; stipules persistent, interpetiolar, triangular, often aristate to spinescent. Inflorescences terminal on main stems and/or axillary short shoots (and then appearing axillary), capitate to congested-fasciculate or -cymose and several flowered or sometimes reduced to 1 flower, bracteate with bracts often fused in pairs. Flowers sessile to shortly pedicellate, bisexual, distylous. Calyx limb (4 or)5(or 6)-lobed. Corolla white to pink or purple, funnelform, inside glabrous to pubescent; lobes (4 or)5(or 6), valvate or valvate-induplicate in bud. Stamens (4 or)5(or 6), inserted in corolla throat, exserted or included; filaments short; anthers dorsifixed. Ovary 5-celled, ovules 1 in each cell, basal, erect, anatropous; stigmas (2 or)3-5(or 6), linear, exserted or included. Fruit capsular, ellipsoid to ovoid, opening through an apical operculum then splitting longitudinally into 5 valves, cartilaginous to woody, calyx limb persistent; aril reticulate, free or adherent to seed; seeds few, medium-sized, ellipsoid, straight, with thin testa; cotyledons rounded; radicle short, inferior.

About 40 species: Himalaya to Japan; 34 species (30 endemic) in China.

1 Aril free from testa (fruit of L. brevisepala and L. dielsiana not seen)   (2)
+ Aril adherent to testa   (18)
2 (1) Bracteoles free   (3)
+ Bracteoles connate   (4)
3 (2) Stipules compressed triangular, shortly acuminate; bracteole longer than calyx.   5 L. dielsiana
+ Stipules long triangular, acuminate; bracteole shorter than calyx or subequal in length.   26 L. scissa
4 (2) Bracteoles connate from base to middle   (5)
+ Bracteoles 1/2-2/3 connate   (10)
5 (4) Bracteoles longer than calyces or nearly as long as calyces   (6)
+ Bracteoles shorter than calyces   (8)
6 (5) Flowers mostly terminal on branches or laterally arranged; calyx lobes lanceolate-subulate, acuminate or shortly acuminate, glabrous; corolla funnelform, lobes not incurved at apex; stamens inserted in corolla tube throat.   8 L. glomerata
+ Flowers 3 terminal on branches; calyx lobes short or long triangular or subovate, very acute, ciliate; corolla narrowly funnelform, lobes incurved at apex; stamens inserted below corolla tube throat   (7)
7 (6) Calyx lobes short or long triangular.   14 L. limprichtii
+ Calyx lobes subovate.   21 L. pumila
8 (5) Leaves thickly papery, glabrous; stipules subtriangular; calyx lobes not ciliate.   2 L. brevisepala
+ Leaves papery, pilose; stipules broadly triangular, glandular on margins; calyx lobes ciliate   (9)
9 (8) Stipules with stiff acicular apiculus, sometimes with stipitate glands on margins; corolla lobes triangular-oblate, apex obtuse; stamens inserted in corolla tube throat.   29 L. velutiniflora
+ Stipules with acicular apiculus, with 2 glandular teeth; corolla lobes ovate-triangular, apex reflexed; stamens inserted below corolla tube throat.   20 L. potaninii
10 (4) Bracteoles equal or subequal in length to calyces; corolla lobes inflexed at apex   (11)
+ Bracteoles longer than calyces; corolla lobes not inflexed at apex   (13)
11 (10) Flowers with pedicels, 3-9 mm.   28 L. umbellata
+ Flowers sessile or subsessile   (12)
12 (11) Corolla small, 11-14 mm; free portion of bracteole triangular.   16 L. oblonga
+ Corolla large, to ca. 16 mm; free portion of bracteole broadly triangular.   9 L. gracilis
13 (10) Stigmas 3-lobed, filiform   (14)
+ Stigmas 5-lobed, linear   (15)
14 (13) Leaves thickly papery; bracteoles sparsely ciliate; calyx lobes shortly acuminate; corolla purplish red, lobes 4 or 5, ovate-lanceolate; stamens inserted above corolla tube throat.   17 L. ordosica
+ Leaves leathery; bracteoles pilose; calyx lobes obtuse; corolla white, lobes 5, sublanceolate; stamens inserted below corolla tube throat.   24 L. scabrida
15 (13) Leaves papery, 3-10 cm, petioles ca. 24 mm; calyx lobes triangular-ovate; corolla lobes with 3 obtuse teeth, incurved.   12 L. kumaonensis
+ Leaves subleathery or leathery, to 4 cm, petioles 4-7 mm; calyx lobes broadly oblong to suborbicular; corolla lobes obtuse   (16)
16 (15) Leaves ovate to broadly lanceolate, whitish or pale yellow abaxially when dry, veinlets indistinct abaxially.   4 L. coriaceifolia
+ Leaves ovate, ovate-oblong, elliptic, or suborbicular, pale green or iron-gray abaxially when dry, veinlets conspicuous abaxially   (17)
17 (16) Flowers 3-7 fascicled, terminal or in axils of upper 3-6 pairs of leaves, subsessile or pedicels 0.3-2.6 mm.   30 L. vestita
+ Flowers (1-)3-5 fascicled, terminal on ultimate branches of inflorescence, forming a thyrse, pedicels to 8 mm.   33 L. yangshuoensis
18 (1) Bracteoles separated, scaly.   1 L. beichuanensis
+ Bracteoles connate   (19)
19 (18) Bracteoles nearly as long as corolla   (20)
+ Bracteoles obviously longer or shorter than calyces   (24)
20 (19) Stigmas 2-lobed.   13 L. lanata
+ Stigmas 5-lobed   (21)
21 (20) Bracteoles tomentose   (22)
+ Bracteoles pubescent or ciliate on upper portion   (23)
22 (21) Bracteoles slightly longer than calyces; calyx lobes ovate, acuminate.   10 L. handeliana
+ Bracteoles shorter than calyces; calyx lobes ovate-oblong, slightly obtuse.   27 L. tomentella
23 (21) Leaves 1-4 × ca. 2 cm, petioles to 1 cm; bracteoles subulate-acuminate at apex.   23 L. rehderiana
+ Leaves 4-9 × 1.25-2 mm, petioles very short or subsessile; bracteoles rounded at apex, apiculate.   25 L. schneideri
24 (19) Bracteoles obviously longer than calyces   (25)
+ Bracteoles obviously shorter than calyces   (29)
25 (24) Stigmas usually 3-lobed.   32 L. xizangensis
+ Stigmas usually 5-lobed   (26)
26 (25) Leaves glabrous on both surfaces   (27)
+ Leaves hairy on both surfaces, at least ciliate along margins   (28)
27 (26) Stipules triangular, glabrous; flowers subsessile; calyx lobes densely fimbriate-ciliate; corolla pallid purple, narrowly funnelform, lobes lanceolate; stamens inserted above corolla tube throat.   3 L. buxifolia
+ Stipules ovate-triangular, covered with soft hairs; flowers with pedicel 3-5(-8) mm; calyx lobes shortly ciliate; corolla purplish blue, funnelform, lobes broadly ovate; stamens inserted below corolla tube throat.   6 L. diffusa
28 (26) Stipules compressed triangular or triangular, acuminate, apiculus rarely with 1 or 2 lateral glandular teeth; calyx lobes acuminate.   10 L. handeliana
+ Stipules broadly triangular, cuspidate-apiculate; calyx lobes obtuse to truncate.   19 L. pilosa
29 (24) Stigmas more than 3-lobed   (30)
+ Stigmas 2- or 3-lobed   (32)
30 (29) Corolla lobes subovate, apex inflexed and beaked.   11 L. hirsutiflora
+ Corolla lobes ovoid-lanceolate, apex mucronate or slightly obtuse   (31)
31 (30) Leaves slightly thick; bracteoles abruptly acuminate at apex; calyx lobes triangular-lanceolate; corolla lobes acute, margins revolute.   18 L. parvifolia
+ Leaves papery, margins obviously recurved; bracteoles abruptly acute; calyx lobes oblong-ovate; corolla lobes slightly obtuse.   22 L. purdomii
32 (29) Corolla broadly funnelform; stamens inserted below corolla tube throat   (33)
+ Corolla funnelform; stamens inserted in corolla tube throat   (34)
33 (32) Stipules apiculate at apex, sparsely pilose at base; bracteole lobes triangular; corolla white, pallid pink, or purple, lobes apically inflexed and beaked.   15 L. ludlowii
+ Stipules lacerate at apex, with glandular hairs along margins; bracteole lobes lanceolate or ensiform-lanceolate; corolla purple, lobes shallowly 3-parted at apex, margins wavy.   34 L. yui
34 (32) Stipules with fimbriate collaters along margins; corolla pallid blue or purplish or purplish red; stigmas 2-lobed.   7 L. forrestii
+ Stipules usually glandular along margins; corolla white or pallid red, stigmas 3-lobed.   31 L. wilsonii

  • List of lower taxa


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