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FOC | Family List | FOC Vol. 22 | Poaceae

56. Melica Linnaeus, Sp. Pl. 1: 66. 1753.

臭草属 chou cao shu

Perennial, tufted or rhizomatous. Culms erect or ascending. Leaf sheaths with fused margins; leaf blades linear; ligule membranous, often cylindrical and then sometimes with lobe on side opposite blade. Panicle spreading or more often contracted, sometimes scanty or racemelike; pedicels drooping, pubescent below spikelet. Spikelets weakly laterally compressed, composed of 1–3 lower fertile florets and a few upper reduced florets, these often compressed into a terminal cluster of rudimentary scales, disarticulating below lowest floret, tardily between florets, or spikelet falling entire; glumes well developed, broadly lanceolate or ovate, often not keeled, membranous or papery, equal or lower glume shorter, 1–5-veined, apex obtuse or acute; floret callus small, glabrous, obtuse; lemmas broadly lanceolate or ovate, usually herbaceous, sometimes largely membranous, back rounded, smooth, scabrid or hairy; 5–9(–13)-veined, apex membranous, obtuse, acute, or shallowly 2-lobed; palea usually shorter than lemma, or as long as lemma in upper florets, keels scabrid or ciliolate. Stamens 3. x = 9.

About 90 species:temperate and subtropical regions of the world, except Australia; 23 species (eight endemic) in China.


1 Lowest lemma conspicuously long-hairy from base to apex, on marginal or all veins, hairs 2–5 mm   (2)
+ Lowest lemma glabrous or with short scattered hairs   (4)
       
2 (1) Lemmas long-pilose on all veins.   1 M. persica
+ Lemmas long-ciliate on marginal veins only   (3)
       
3 (2) Panicle rather lax, often 1-sided, central axis usually visible; all leaf sheaths antrorsely scaberulous; leaf blades rolled; lower glume 2/3–4/5 length of upper.   2 M. ciliata
+ Panicle dense, cylindrical, central axis not visible; lowermost leaf sheaths retrorsely scabrid or pubescent; leaf blades flat; lower glume 1/2–2/3 length of upper.   3 M. transsilvanica
       
4 (1) Spikelets terminating in 1 sterile lemma, resembling the fertile lemmas but smaller; panicle branches up to 15 cm, often spreading, branchlets present   (5)
+ Spikelets terminating in a globular cluster of rudimentary lemmas; panicle branches usually less than 5 cm, erect, unbranched or almost so, panicle sometimes racemelike (rarely branches longer or branchlets present)   (11)
       
5 (4) Lowest lemma 6–9 mm; leaf sheaths harshly scabrid.   4 M. scaberrima
+ Lowest lemma 4–6.5 mm; leaf sheaths smooth, scabrid or pubescent   (6)
       
6 (5) Plants with slender rhizomes; panicle narrow, branches erect or ascending   (7)
+ Plants lacking slender rhizomes; panicle broad at maturity, branches spreading   (8)
       
7 (6) Ligule 3–5 mm; leaf blades 2–2.5 mm wide; anthers ca. 2 mm.   7 M. longiligulata
+ Ligule 0.3 mm or shorter; leaf blades 2–6 mm wide; anthers 0.5–1 mm.   8 M. przewalskyi
       
8 (6) Culms up to 150 cm tall; leaf blades 6–14 mm wide; ligule 0.2–0.5 mm.   5 M. onoei
+ Culms up to 80 cm tall (if taller, ligule 1–4 mm); leaf blades 2–6 mm wide   (9)
       
9 (8) Panicle with many branches and spikelets; spikelets with purple glumes and green florets; fertile florets 2 or 3.   6 M. schuetzeana
+ Panicle with distant divaricate branches and widely spaced spikelets; spikelets green or gray-green; fertile florets 1 or 2   (10)
       
10 (9) Glumes hyaline, shining, upper glume 6–8 mm; pedicels up to 20 mm; anthers 1.8–2 mm.   9 M. yajiangensis
+ Glumes not conspicuously hyaline and shining, upper glume 5–6 mm; pedicels 3–5 mm; anthers 1–1.25 mm.   10 M. taylorii
       
11 (4) Panicle racemelike, unbranched; spikelets few (3–15), borne directly on main axis   (12)
+ Panicle branched; spikelets more than 15, borne on panicle branches   (14)
       
12 (11) Spikelets 5–8 mm; glumes purplish red; panicle eventually nodding.   11 M. nutans
+ Spikelets 7–10 mm; glumes usually green; panicle erect   (13)
       
13 (12) Glumes subequal, ovate, obtuse.   12 M. grandiflora
+ Glumes unequal, lanceolate, acute.   13 M. pappiana
       
14 (11) Culms up to 1.5 m tall; spikelets 8–14 mm; glumes obviously 5–7-veined   (15)
+ Culms less than 1 m tall; spikelets 3.5–8(–11) mm; glumes with short inconspicuous lateral veins   (16)
       
15 (14) Panicle open, ovate, branches spreading; lemmas hispid below middle.   14 M. turczaninowiana
+ Panicle dense, linear-oblong, branches erect; lemmas glabrous.   15 M. altissima
       
16 (14) Glumes unequal, much shorter than florets; lower glume ca. 1/2 length of adjacent lemma.   16 M. virgata
+ Glumes subequal, large; lower glume slightly shorter to longer than adjacent lemma   (17)
       
17 (16) Lemma apex obtuse or acute   (18)
+ Lemma apex very broad, denticulate-erose, usually 2-lobed or emarginate   (20)
       
18 (17) Panicle with many crowded spikelets, 20–50 on lower branches (including branchlets); leaf blades flat, 2–7 mm wide.   17 M. scabrosa
+ Panicle with few spikelets, 1–6 on lower branches (branchlets absent); leaf blades usually rolled, 1–3 mm wide   (19)
       
19 (18) Plant tufted; ligules ca. 0.5 mm; lemmas coarsely tubercular-scabrid.   18 M. radula
+ Plant rhizomatous; ligule 2–5 mm; lemmas scaberulous.   19 M. secunda
       
20 (17) Panicle lax; ligules with ca. 3 mm lobe on side opposite blade; anthers 1.2–2.2 mm.   20 M. kozlovii
+ Panicle very dense; ligules without lobe on side opposite blade; anthers 0.6–1 mm   (21)
       
21 (20) Spikelets silvery green; lemmas narrowly membranous at apex; ligule abaxially glabrous.   21 M. tangutorum
+ Spikelets purple or yellow; lemmas membranous and slightly enlarged in upper 1/3; ligule abaxially pubescent   (22)
       
22 (21) Panicle purple; spikelets 5–8 mm; ligules 0.8–1.5 mm.   22 M. tibetica
+ Panicle yellow; spikelets 8–11 mm; ligules 2–4 mm.   23 M. subflava

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