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FOC | Family List | FOC Vol. 17 | Lamiaceae

54. Microtoena Prain, Hooker's Icon. Pl. 19: t. 1872. 1889.

冠唇花属 guan chun hua shu

Herbs erect. Leaf blade margin dentate. Cymes dichotomous, axillary and/or in terminal panicles; bracts and bracteoles linear, early deciduous. Calyx campanulate, 10-veined; teeth 5, triangular to linear, subequal or posterior teeth much longer; fruiting calyx dilated or saccate-dilated, base rounded. Corolla yellow, rarely white, marked with purple-red or brown on upper lip; tube exserted, straight, base narrow, dilated above middle, annulus absent inside; upper lip straight, galeate, base obliquely cuneate or sometimes short clawed; lower lip spreading, 3-lobed, middle lobe ligulate to ovate, lateral lobes ovate or circular. Stamens 4, subequal, included in upper lip or slightly exserted; filaments complanate, usually glabrous; anther cells 2, apically divaricate, becoming confluent. Style as long as stamens, anterior lobe subulate, posterior lobe short to inconspicuous. Nutlets ovoid, glabrous, adaxially veined, abaxially rounded, constricted at base, areole small.

About 24 species: Asia, 20 species in China.

1 Middle lobe of lower corolla lip narrow, ligulate; upper corolla lip always purple-red or brown.   (2)
+ Middle lobe of lower lip of corolla circular to ovate or obovate; upper corolla lip yellow (yellow and purple-red in M. delavayi).   (5)
2 (1) Dichotomous cymes 2- or 3-flowered, axillary   2 Microtoena pauciflora
+ Dichotomous cymes many flowered, axillary or in terminal panicles.   (3)
3 (2) Leaf base broadly truncate-cuneate, decurrent into winged petiole 5. M. insuavis   5 Microtoena insuavis
+ Leaf base cordate.   (4)
4 (3) Calyx ca. 3 mm; leaves triangular-ovate, rarely oblong-ovate; stem spreading villous, retrorsely minutely tomentose   1 Microtoena patchoulii
+ Calyx 6.5 mm; leaves ovate-cordate; stem sparsely glandular villous, spreading hispid   3 Microtoena mollis
5 (1) Cymes ± sessile, in terminal ± spicate panicles   4 Microtoena subspicata
+ Cymes pedunculate and/or axillary, not in terminal ± spicate panicle.   (6)
6 (5) Calyx teeth triangular-subulate, apex slightly curved or hooked.   (7)
+ Calyx teeth triangular to linear, if triangular-subulate and apically slightly curved, then posterior tooth linear, long.   (11)
7 (6) Posterior tooth of calyx ca. 2 × as long as other teeth   16 Microtoena urticifolia
+ Calyx teeth subequal, or posterior tooth only slightly longer.   (8)
8 (7) Panicle compact; calyx glabrous except for ciliolate margins   18 Microtoena prainiana
+ Panicle lax; calyx glandular hairy.   (9)
9 (8) Leaves subglabrous except sometimes for veins; calyx broadly campanulate, finely glandular puberulent outside   20 Microtoena megacalyx
+ Leaves hairy; calyx campanulate.   (10)
10 (9) Leaves ovate-cordate to triangular-ovate, adaxially finely strigose, abaxially densely appressed tawny hispidulous along veins   17 Microtoena moupinensis
+ Leaves oblong-ovate, adaxially sparsely minutely hispid, abaxially spreading hispid along veins   19 Microtoena longisepala
11 (6) Posterior calyx tooth 2.5-3 × as long as other teeth, anterior 4 teeth minute, narrowly triangular to triangular-subulate, equal.   (12)
+ Posterior calyx tooth up to 2 × as long as anterior teeth.   (13)
12 (11) Stems, petioles, and abaxial leaf surface densely hairy; cymes in short compact panicles; posterior calyx tooth linear, ca. 3 × as long as other teeth; upper corolla lip not auriculate   13 Microtoena miyiensis
+ Stem, petioles, and abaxial leaf surfaces sparsely hairy; cymes in elongated loose panicles; posterior calyx tooth lanceolate ca. 2.5 × as long as other teeth; upper corolla lip basally ligulate-auriculate   12 Microtoena maireana
13 (11) Calyx teeth lanceolate to linear-lanceolate.   (14)
+ Calyx teeth triangular-lanceolate, triangular, or oblong.   (17)
14 (13) Cymes 1-3-flowered, axillary.   (15)
+ Cymes many flowered, in terminal panicles though sometimes also axillary.   (16)
15 (14) Cymes 2- or 3-flowered; peduncle 3-5 mm   11 Microtoena albescens
+ Cymes 1-flowered; peduncle 6-15 mm   10 Microtoena affinis
16 (14) Upper corolla lip often purple-red (yellow in var. lutea); terminal panicles ca. 2-3 × 3-41 cm   6 Microtoena delavayi
+ Corolla uniformly yellow; terminal panicles 4.5-13 × ca. 3.5 cm   9 Microtoena stenocalyx
17 (13) Calyx teeth only slightly unequal, posterior tooth longer, oblong, apex subacute   14 Microtoena robusta
+ Calyx teeth obviously unequal, posterior tooth much longer, triangular-ovate to triangular-lanceolate.   (18)
18 (17) Calyx teeth triangular-ovate; panicle of 3(-5)-flowered cymes, terminal on lateral branches, or a few cymes solitary, nearly apical   15 Microtoena vanchingshanensis
+ Calyx teeth triangular-lanceolate; inflorescences not as above.   (19)
19 (18) Cymes axillary and terminal, lax, branches 1-sided; at least some peduncles over 2 cm; bracts reduced, minute, or absent   7 Microtoena muliensis
+ Cymes all dichotomous, axillary; peduncles (1-)1.6 cm; bracts lanceolate, 2-3 mm   8 Microtoena omeiensis

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