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72. Millettia Wight & Arnott, Prodr. Fl. Ind. Orient. 263. 1834.

崖豆藤属 ya dou teng shu

Authors: Zhi Wei & Les Pedley

Trees, shrubs, or lianas. Stipules present, persistent or caducous. Leaves imparipinnate; stipels present or absent; leaflet blades opposite or rarely subalternate. Inflorescence a pseudoraceme, unbranched (but branched in Millettia ichthyochtona), rachis with lateral flowering branchlets reduced to knobs or short spur at each node, flowers 2-5 in a clusters or fascicles per node; bracts and bracteoles present. Calyx tube broad, shortly 5-toothed; adaxial 2 teeth ± connate. Corolla standard large, with or without basal cal­luses. Vexillary stamen free in bud sometimes and becoming adherent to other 9 when mature. Ovary linear; style straight or in­flexed. Legume 2-valved, flat, sometimes cylindric or inflated only around ellipsoid seeds, tardily dehiscent, not winged or with 2 wings perpendicular to abaxial or both sutures. Seeds 1 to several per legume, lenticular or globose; funicle fleshy, swelling at adaxi­al side, surrounding and clasping hilum as a short ringlike aril.

About 100 species: subtropical and tropical Africa, Asia, and Australia; 18 species (six endemic) in China.

Many species are used in China as medicinal drugs, insecticide, or fish poison. A few species are cultivated as ornamentals.


1 Petals outside glabrous (but not always explicitly stated in description; corolla unknown in M. entadoides)   (2)
+ Petals outside pubescent   (8)
       
2 (1) Leaf rachis and leaflet blades glabrous   (3)
+ Leaf rachis and leaflet blades at least abaxially with trichomes   (5)
       
3 (2) Leaflets subalternate, blades obliquely ovate, base asymmetric; legume stipitate.   1 M. ichthyochtona
+ Leaflets opposite, blades oblong-lanceolate, oblanceolate, or oblong, base symmetric; legume sessile   (4)
       
4 (3) Leaflet blades 12-25 cm, secondary veins 11-13 on each side of midvein; seeds thickly lenticular; trees.   3 M. leptobotrya
+ Leaflet blades 9-12 cm, secondary veins 9-12 on each side of midvein; seeds reniform; lianas.   13 M. entadoides
       
5 (2) Leaflets 13-17, blades abaxially brown sericeous; legume inflated, rough with dense warts; lianas.   12 M. pachycarpa
+ Leaflets 7-11(or 13), blades abaxially yellow tomentose, scarlet strigose on midvein and margins, or pilose on midvein; legume flat, without warts; trees   (6)
       
6 (5) Leaflet blades abaxially yellow tomentose; legume sutures both with 2 ca. 1 cm wide woody wings.   2 M. tetraptera
+ Leaflet blades abaxially scarlet strigose on midvein and margins or pilose on midvein; legume sutures both only thickened   (7)
       
7 (6) Leaflet blades abaxially scarlet strigose on midvein and margins; pseudoracemes erect.   4 M. erythrocalyx
+ Leaflet blades abaxially pilose on midvein; pseudoracemes pendulous.   5 M. leucantha
       
8 (1) Stipels absent   (9)
+ Stipels present   (11)
       
9 (8) Leaflet blades broadly elliptic, membranous, both surfaces pubescent, apex acute; legume flat.   6 M. pubinervis
+ Leaflet blades oblong-elliptic to lanceolate-elliptic or lanceolate to oblong-lanceolate, glabrous or sparsely pilose adaxially, apex caudate; legume inflated   (10)
       
10 (9) Leaflet blades oblong-elliptic to lanceolate-elliptic, papery, abaxially brown tomentose, adaxially shiny and sparsely pilose.   11 M. sapindifolia
+ Leaflet blades lanceolate to oblong-lanceolate, subleathery, abaxially sparsely strigose and adaxially glabrous.   7 M. austroyunnanensis
       
11 (8) Leaflets 13-19   (12)
+ Leaflets (5 or)7-13(or 15)   (14)
       
12 (11) Stipules ca. 8 mm, persistent; branchlets reddish brown tomentose.   8 M. cubittii
+ Stipules 1-3 mm, caducous; branchlets yellow tomentose or grayish yellow pubescent   (13)
       
13 (12) Pseudoracemes 20-25 cm; flowers 1.3-1.6 cm; corolla white or lilac; legume tomentose; trees 8-15 m tall.   16 M. velutina
+ Pseudoracemes 6-15 cm; flowers 0.9-1.2 cm; corolla pale red to scarlet; legume gray pubescent but glabrescent; erect shrubs or trees 3-8 m tall.   17 M. pulchra
       
14 (11) Leaf rachis 15-20 cm; leaflet blades 4-5.5 cm.   18 M. oraria
+ Leaf rachis 25-40 cm; leaflet blades 7-18 cm   (15)
       
15 (14) Leaflets 5; stems slender, without lenticels.   9 M. lantsangensis
+ Leaflets 7-13; stems thick, with scattered lenticels   (16)
       
16 (15) Leaflet blade secondary veins 13-17 on each side of midvein; lianas.   10 M. pachyloba
+ Leaflet blade secondary veins 7-10 on each side of midvein; trees or erect shrubs   (17)
       
17 (16) Leaflet blades abaxially glabrescent; inflorescences longer than subtending leaf; flowers ca. 1.8 cm; trees.   14 M. macrostachya
+ Leaflet blades abaxially densely brown pubescent especially on veins; inflorescences shorter than subtending leaf; flowers ca. 1 cm; erect shrubs.   15 M. griffithii

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