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72. Mosla (Bentham) Buchanan-Hamilton ex Maximowicz, Bull. Acad. Imp. Sci. Saint-Pétersbourg. sér. 3, 20: 456. 1875.

石荠苎属 shi qi zhu shu

Hedeoma Persoon sect. Mosla Bentham, Labiat. Gen. Spec. 366. 1834.

Herbs aromatic, annual. Leaves petiolate, dentate, abaxially conspicuously impressed glandular. Verticillasters 2-flowered, in terminal racemes; bracts small or lower ones leaflike. Flowers pedicellate. Calyx campanulate, 10-veined, throat hairy, limb subequally 5-toothed or 2-lipped; upper lip 3-toothed, teeth acute to obtuse; lower lip 2-toothed, teeth lanceolate; fruiting calyx dilated, with base swollen on 1 side. Corolla white or rose to purple-red, tube glabrous or hairy annulate inside; limb nearly 2-lipped, upper lip emarginate; lower lip 3-lobed, margin crenate; lateral lobes smaller than middle lobe, emarginate. Stamens 4, posterior 2 fertile, anterior 2 reduced, with indistinct anther cells; anther cells 2, divaricate. Style apex subequally to equally 2-cleft. Nutlets subglobose to globose, sparsely netted or pitted with small depressions; areolae basal, dotlike.

About 22 species: Cambodia, China, India, Indonesia, Japan, Korea, Laos, Malaysia, Philippines, Vietnam; 12 species in China.


1 Bracts ovate-lanceolate to circular-obovate; calyx subequally 5-toothed; nutlets pitted with small depressions.   (2)
+ Bracts ovate-lanceolate, lanceolate, to needlelike, generally shorter than flowers; calyx 2-lipped; nutlets sparsely netted rarely pitted with small depressions.   (6)
       
2 (1) Bracts ovate-lanceolate, 4-6(-9) mm.   (3)
+ Bracts broadly ovate, subcircular, to circular-obovate, 1.5-7 mm.   (4)
       
3 (2) Racemes dense, occupying a large part of each stem or branch; leaves few   4 Mosla longispica
+ Racemes lax, occupying a small part of each stem or branch; leaves many   5 Mosla pauciflora
       
4 (2) Bracts 1.5-2.5 mm, loosely arranged   3 Mosla soochowensis
+ Bracts 4-7 × 3-5 mm, overlapping.   (5)
       
5 (4) Leaves lanceolate, 5-13 mm; corolla ca. 10 mm   2 Mosla hangchowensis
+ Leaves linear-oblong to linear-lanceolate, 2-4(-7) mm; corolla ca. 5 mm   1 Mosla chinensis
       
6 (1) Upper calyx teeth acute.   (7)
+ Upper calyx teeth obtuse.   (8)
       
7 (6) Stem retrorse fine puberulent, glabrescent; leaves coarsely 3-5-serrate   7 Mosla grosseserrata
+ Stem densely pubescent; leaves serrate   6 Mosla scabra
       
8 (6) Plants leafless; stem retrorse gray pubescent; each branch with a many-flowered raceme; bracts ovate-lanceolate, 2.5-3.5 mm, 1.5-2 × as long as pedicels   8 Mosla exfoliata
+ Plants many leaved.   (9)
       
9 (8) Corolla ca. 2.5 mm; plants sparsely villous, puberulent; leaves ovate to ovate-lanceolate   9 Mosla cavaleriei
+ Corolla 4-10 mm or more; plants not sparsely villous.   (10)
       
10 (9) Leaves remotely serrate; bracts needlelike to linear-lanceolate   10 Mosla dianthera
+ Leaves crenate to crenate-serrate; bracts ovate to lanceolate.   (11)
       
11 (10) Bracts much longer than pedicels; calyx ca. 2.7 mm; corolla slightly longer than calyx   11 Mosla longibracteata
+ Bracts as long as pedicels; calyx less than 2 mm; corolla ca. 3 × as long as calyx   12 Mosla formosana

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