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FOC | Family List | FOC Vol. 13 | Nyssaceae

3. Nyssa Linnaeus, Sp. Pl. 2: 1058. 1753.

蓝果树属 lan guo shu shu

Trees dioecious. Leaves often crowded near ends of branches, estipulate. Flowers unisexual, usually in heads or short racemes, in axil of a bract with 2 bracteoles. Male flowers 5-merous. Stamens 10, arranged in two alternate whorls; filaments linear; anthers 2-celled, dorsifixed, with lateral lengthwise slits; disk pulvinate. Female flowers (4 or)5-merous; staminodes usually present. Ovary inferior, 1(or 2)-loculed, 1-ovuled; style bifid, with stigmatic tissue at inside of stylar arms. Fruit drupaceous, ± laterally flattened, with persistent calyx and disk. n = 22 [in Nyssa javanica (Blume) Wangerin (Mehra & Bawainin, Evolution 23: 466-481. 1969)].

About 12 species: four in North America, one in Costa Rica, one from India to Malaysia, and seven (five endemic) in China.

According to C. Q. Zhang (pers. comm.), most Chinese species of Nyssa are endangered and some of them may already have become extinct.

1 Flowers pedicellate, in umbels or racemes   (2)
+ Flowers in heads, sessile or male ones shortly pedicellate   (3)
2 (1) Branchlets, petioles, and pedicels pilose when young, subglabrous when old.   5 N. sinensis
+ Branchlets, petioles, and pedicels persistently pilose.   4 N. shweliensis
3 (1) Leaves thin, papery   (4)
+ Leaves thick, leathery   (5)
4 (3) Leaf blade subelliptic, to 5 × 3.5 cm, petiole to 1 cm.   2 N. leptophylla
+ Leaf blade suboblong, not less than 8 × 4 cm, petiole not less than 1.5 cm.   6 N. wenshanensis
5 (3) Drupes usually less than 1.2 cm; branchlets, pedicels, and leaves glabrous.   3 N. shangszeensis
+ Drupes not less than 1.4 cm; branchlets, pedicels, and leaves pilose at least when young   (6)
6 (5) Branchlets, pedicels, and leaves abaxially tomentose when young, subglabrous when mature.   1 N. javanica
+ Branchlets, pedicels, and leaves abaxially persistently pilose.   7 N. yunnanensis

Lower Taxa


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