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9. Paris Linnaeus, Sp. Pl. 1: 367. 1753.
重楼属 chong lou shu
Liang Songyun (梁松筠 Liang Song-jun); Victor G. Soukup1
Daiswa Rafinesque; Euthyra Salisbury; Kinugasa Tatewaki & Suto.
Herbs perennial. Rhizome slender or thickened. Stem erect, simple. Leaves 4 to many, very rarely 3, in a terminal whorl, petiolate, lanceolate to ovate, with 3 main veins and anastomosing veinlets. Flowers bisexual, solitary, terminal, pedunculate. Tepals 3--8, in 2 whorls, free; outer ones green, rarely ± white, ovate to lanceolate; inner ones linear or occasionally absent. Stamens 8--24 or more, 2--8 × as many as tepals; filaments narrow, flat; anthers basifixed, often with convex connective apically. Ovary subglobose, 1-loculed with parietal placentation or 4--10-loculed with axile placentation. Style short; stigma lobes 4--10. Fruit a berry or a berrylike capsule, indehiscent or loculicidal, several to many seeded.
About 24 species: Bhutan, China, India, Japan, Korea, Laos, Mongolia, Myanmar, Nepal, Russia, Sikkim, Thailand, Vietnam; Europe; 22 species (12 endemic) in China.
The rhizomes of many species are used medicinally in China.
Also see JI Yun-Heng, ZHOU Zhe-Kun & LI Heng. 2007. Four new synonyms in the genus Paris (Trilliaceae. Acta Phytotax. Sin. 45: 388-390.
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1 |
Ovary 4--10-loculed, with axile placentation; berry indehiscent; seeds sometimes with a spongy (never succulent) aril on 1 side. |
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(2) |
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Ovary 1-loculed with parietal placentation; berrylike capsule dehiscing irregularly; seeds with a succulent aril. |
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(9) |
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2 (1) |
Rhizome creeping, elongate, slender, 2--5 mm thick; seeds without aril, funicle not swollen. |
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(3) |
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Rhizome 8--45 mm thick; seeds ± enveloped by whitish green, spongy aril formed by swollen funicle. |
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(5) |
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3 (2) |
Leaves usually 6--8; outer tepals oblong-lanceolate to ovate-lanceolate, 10--25(--30) mm wide |
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20 Paris verticillata |
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Leaves usually 4; outer tepals narrowly lanceolate, 3--9 mm wide. |
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(4) |
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4 (3) |
Leaves oblong-lanceolate or ovate-elliptic; outer tepals reflexed, 3--5 mm wide |
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21 Paris bashanensis |
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Leaves broadly obovate or ovate; outer tepals spreading horizontally, 5--9 mm wide |
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22 Paris quadrifolia |
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5 (2) |
Leaves 9--10 |
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19 Paris dulongensis |
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Leaves 4--6. |
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(6) |
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6 (5) |
Leaves wrinkled, obovate, 10--15 × 6.3--10 cm, basally rounded |
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18 Paris rugosa |
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Leaves not wrinkled. |
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(7) |
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7 (6) |
Leaves elliptic or oblanceolate, base subcuneate |
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17 Paris vaniotii |
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Leaves oblong, ovate- or obovate-oblong, or oblong-lanceolate, base shallowly cordate or rounded. |
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(8) |
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8 (7) |
Seeds white or yellow-red, slightly enveloped by swollen funicle |
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16 Paris forrestii |
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Seeds yellow-brown, 1/2 enveloped by greenish white or rarely orange, spongy aril formed by enlarged funicle |
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15 Paris axialis |
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9 (1) |
Stamens 4--6 × as many as outer tepals; seeds enveloped by orange aril |
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1 Paris dunniana |
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Stamens 2 or 3 × as many as outer tepals; seeds enveloped by red aril. |
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(10) |
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10 (9) |
Leaves with white, yellowish, or purple markings. |
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(11) |
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Leaves without markings. |
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(13) |
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11 (10) |
Leaves with purple markings; stamens ca. 3 × as many as leaves |
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12 Paris polyandra |
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Leaves with white or yellowish markings; stamens 2 × as many as leaves. |
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(12) |
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12 (11) |
Outer tepals as many as leaves, shorter than inner ones |
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10 Paris luquanensis |
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Outer tepals fewer than leaves, longer than inner ones |
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11 Paris marmorata |
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13 (10) |
Plants pubescent or papillose-pubescent. |
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(14) |
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Plants glabrous or nearly so. |
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(15) |
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14 (13) |
Free portion of connective 0.5--1 mm |
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8 Paris mairei |
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Free portion of connective 13--15 mm |
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9 Paris wenxianensis |
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15 (13) |
Free portion of connective 6--15 mm. |
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(16) |
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Free portion of connective 1--4(--5) mm (except in P. polyphylla var. pseudothibetica). |
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(17) |
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16 (15) |
Inner tepals pendulous, margin undulate |
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7 Paris undulata |
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Inner tepals (if present) not pendulous, margin not undulate |
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14 Paris thibetica |
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17 (15) |
Filaments ca. 1 mm, shorter than free portion of connective |
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5 Paris daliensis |
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Filaments more than 2 mm, longer than free portion of connective. |
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(18) |
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18 (17) |
Free portion of connective transversely ellipsoid, subglobose, or shortly conical, as wide as or wider than long |
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13 Paris fargesii |
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Free portion of connective inconspicuous or longer than wide. |
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(19) |
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19 (18) |
Leaves (18--)23--26 × (11--)14--16 cm |
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3 Paris vietnamensis |
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Leaves 5--17 × 2--11 cm. |
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(20) |
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20 (19) |
Leaves ovate |
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2 Paris cronquistii |
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Leaves oblong to lanceolate. |
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(21) |
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21 (20) |
Inner tepals dark purple, usually much shorter than outer ones |
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4 Paris delavayi |
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Inner tepals yellow-green, usually slightly shorter or longer than outer ones |
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6 Paris polyphylla |
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List of lower taxa
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