1 |
Sterile leaves monomorphic, spirally arranged, linear or linear-lanceolate, with long white apical seta |
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(2) |
+ |
Sterile leaves dimorphic, arranged in 4 ranks, 2 dorsal and 2 ventral leaves, rarely nearly monomorphic, without long white apical seta |
|
(4) |
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2 (1) |
Leaves thickly leathery, obtuse at apex. |
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1 S. sibirica |
+ |
Leaves thinly leathery, acuminate at apex |
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(3) |
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3 (2) |
Apical seta ca. 1/3 as long as leaves. |
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2 S. vardei |
+ |
Apical seta ca. 1/5 as long as leaves. |
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3 S. indica |
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4 (1) |
Stems and branches cylindrical, often reddish; sterile leaves nearly monomorphic, adpressed to stems and branches |
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(5) |
+ |
Branches often compressed; leaves dimorphic, dorsal leaves ascending, ventral leaves spreading |
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(6) |
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5 (4) |
Leaves not white-margined. |
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4 S. sanguinolenta |
+ |
Leaves distinctly white-margined. |
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5 S. albocincta |
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6 (4) |
Strobili lax, not distinct from sterile stems; sporophylls monomorphic or dimorphic, almost conform with sterile leaves at same rank (non-resupinate) (see also 42b) |
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(7) |
+ |
Strobili compact, distinct from sterile stems; sporophylls in 4 ranks, monomorphic or dimorphic (dorsal sporophylls larger than ventral ones, rarely smaller) |
|
(9) |
|
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7 (6) |
Strobili cylindrical; sporophylls monomorphic, smaller than ventral leaves. |
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6 S. helvetica |
+ |
Strobili rather loosely dorsiventrally compressed; sporophylls dimorphic, nearly conform with sterile leaves at same rank |
|
(8) |
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8 (7) |
Leaves denticulate at margin. |
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7 S. nipponica |
+ |
Leaves shortly ciliolate at margin. |
|
8 S. pseudonipponica |
|
|
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9 (6) |
Strobili tetragonal; sporophylls nearly monomorphic |
|
(10) |
+ |
Strobili dorsiventrally complanate; dorsal and ventral sporophylls dimorphic |
|
(41) |
|
|
|
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10 (9) |
Stems forming rosettes, curling inward when dry |
|
(11) |
+ |
Stems not forming rosettes, not curling inward when dry |
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(12) |
|
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|
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11 (10) |
Dorsal and ventral leaves denticulate at margin. |
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9 S. tamariscina |
+ |
Dorsal leaves reflexed at margin; acroscopic margin of ventral leaves brown, membranous, lacerate. |
|
10 S. pulvinata |
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12 (10) |
Main stems creeping or prostrate in basal portion to scandent; rhizophores borne throughout stems |
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(13) |
+ |
Main stems erect, suberect, or ascending with short prostrate basal portion; rhizophores usually borne on basal to middle portions of main stem |
|
(24) |
|
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13 (12) |
Main stems scandent, up to 1-2 m or longer |
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(14) |
+ |
Main stems creeping, decumbent, or scandent, usually less than 1 m |
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(15) |
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14 (13) |
Axillary and ventral leaves with large basal auricles; dorsal leaves slightly white-margined; sporophylls ovate, acute or cuspidate; base of rhizophores without spines. |
|
11 S. willdenowii |
+ |
Axillary and ventral leaves with small basal auricles; dorsal leaves white-margined; sporophylls ovate-lanceolate, acuminate; base of rhizophores with a few spines. |
|
12 S. helferi |
|
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15 (13) |
Plants xerophytic, curling when dry; leaves ciliolate at margin |
|
(16) |
+ |
Plants not xerophytic, not curling when dry; leaves entire or denticulate at margin |
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(18) |
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16 (15) |
Dorsal leaves slightly smaller than and overlapping ventral leaves; leaves slightly lacerate and densely long ciliolate at margin. |
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13 S. nummularifolia |
+ |
Dorsal leaves much smaller than and not largely overlapping ventral leaves |
|
(17) |
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17 (16) |
Stems and branches scarlet; ventral leaves reflexed; margin of leaves sparsely toothed and long ciliolate at acroscopic base, entire elsewhere. |
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14 S. rossii |
+ |
Stems and branches stramineous; ventral leaves not reflexed; margin of leaves shortly ciliolate throughout. |
|
15 S. sinensis |
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18 (15) |
Leaves entire |
|
(19) |
+ |
Leaves denticulate or ciliolate at margin |
|
(21) |
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19 (18) |
Dorsal leaves auriculate at basiscopic base. |
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16 S. limbata |
+ |
Dorsal leaves not auriculate |
|
(20) |
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20 (19) |
Sporophylls not carinate on abaxial surface. |
|
17 S. uncinata |
+ |
Sporophylls carinate on abaxial surface. |
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18 S. hainanensis |
|
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21 (18) |
Stems not articulate; rhizophores borne on ventral side in axils of branches |
|
(22) |
+ |
Stems slightly articulate beneath branching; rhizophores borne on dorsal side in axils of branches |
|
(23) |
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22 (21) |
Leaves not white-margined, leaves on main stems and lateral branches ciliolate at margin, those on ultimate branches often subentire. |
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19 S. siamensis |
+ |
Leaves white-margined, denticulate or shortly ciliolate. |
|
20 S. davidii |
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23 (21) |
Stems with a single vascular bundle. |
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21 S. remotifolia |
+ |
Stems with 3 vascular bundles. |
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22 S. kraussiana |
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24 (12) |
Rhizophores borne on basal and middle portions of main stem or sometimes also on upper portion; leaves on main stem dimorphic |
|
(25) |
+ |
Rhizophores restricted to basal or lower portion of main stem or to prostrate rhizome; leaves on main stem monomorphic |
|
(32) |
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25 (24) |
Plants erect; lateral branches regularly pinnately branched, twice forked, or simple; leaves entire or minutely denticulate at apex |
|
(26) |
+ |
Plants erect, suberect, or ascending with prostrate rhizome; lateral branches many times pinnately branched; leaves denticulate |
|
(28) |
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26 (25) |
Main stems 3-5 mm in diam., apical part black when dry; stomata invisible on dorsal leaves. |
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23 S. picta |
+ |
Main stems 2-3 mm in diam., apical part not black when dry; stomata on adaxial surface of dorsal leaves visible |
|
(27) |
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27 (26) |
Lateral branchlets simple or forked, or basal branchlets often forked; dorsal leaves overlapping distally. |
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24 S. delicatula |
+ |
Lateral branchlets simple, sometimes with irregular longer branches; dorsal leaves parallel, not overlapping distally. |
|
25 S. wallichii |
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28 (25) |
Leaves subleathery, shiny when dry; dorsal leaves aristate, arista ca. 1/2 as long as leaves or longer. |
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26 S. commutata |
+ |
Leaves herbaceous, not shiny when dry; dorsal leaves acuminate to aristate, arista very short |
|
(29) |
|
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29 (28) |
Ventral leaves 4.5-7 mm. |
|
27 S. rolandi-principis |
+ |
Ventral leaves less than 4.5 mm |
|
(30) |
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30 (29) |
Ventral leaves ovate-triangular or ovate-lanceolate; dorsal leaves aristate; surface of dorsal and ventral leaves scabrous. |
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28 S. scabrifolia |
+ |
Ventral leaves ovate-oblong or oblong-falcate; dorsal leaves acuminate or very shortly aristate; surface of leaves glabrous or that of ventral leaves scabrous |
|
(31) |
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31 (30) |
Dorsal and ventral leaves glabrous. |
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29 S. doederleinii |
+ |
Ventral leaves scabrous. |
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30 S. trachyphylla |
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32 (24) |
Stems and branches pubescent |
|
(33) |
+ |
Stems and branches glabrous |
|
(37) |
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33 (32) |
Main stems isotomously branched. |
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31 S. pubescens |
+ |
Main stems pinnately branched |
|
(34) |
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34 (33) |
Plants 45-100 cm high or more; axillary leaves biauriculate at base, ventral leaves auriculate at acroscopic base |
|
(35) |
+ |
Plants 10-45 cm high; axillary leaves not auriculate at base |
|
(36) |
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35 (34) |
Leaf portion of main stems and branches pubescent on both sides; main stems zigzag; leaves white-margined. |
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32 S. trichoclada |
+ |
Leaf portion of main stems glabrous, ventral side of branches pubescent; main stems not zigzag; leaves not white-margined. |
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33 S. pseudopaleifera |
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36 (34) |
Leaves thick, shrunken when dry; leaves on main stems peltate. |
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34 S. braunii |
+ |
Leaves thin, not shrunken; leaves on main stems basifixed. |
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35 S. biformis |
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37 (32) |
Plants usually more than 50 cm high, main stems robust, up to 4-5 mm in diam.; branches including leaves 5-10 mm wide. |
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36 S. superba |
+ |
Plants usually less than 50 cm high, main stems 1-2 mm in diam., not so robust; branches including leaves less than 5 mm wide |
|
(38) |
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38 (37) |
Leaves on main stems approximate |
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(39) |
+ |
Leaves on main stems distant |
|
(40) |
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39 (38) |
Main stems reddish. |
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37 S. stauntoniana |
+ |
Main stems stramineous. |
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38 S. involvens |
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40 (38) |
Leaves on main stems peltate at base, not white-margined, ciliolate, very shrunken when dry. |
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39 S. mairei |
+ |
Leaves on main stems basifixed, white-margined, denticulate, not shrunken when dry. |
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40 S. moellendorffii |
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41 (9) |
Strobili non-resupinate, i.e., dorsal sporophylls smaller than ventral ones |
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(42) |
+ |
Strobili resupinate, i.e., dorsal sporophylls larger than ventral ones |
|
(46) |
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42 (41) |
Strobili rather lax, often forked (see also 6a) |
|
(43) |
+ |
Strobili shortly compact, solitary (or sometimes forked in S. prostrata) |
|
(44) |
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43 (42) |
Fertile branches short, pinnately branched; leaves up to 3.2 ?1.8 mm, margin denticulate, shortly ciliolate at base. |
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41 S. pallidissima |
+ |
Fertile branches erect or ascending, forked; leaves up to 2.3 ?1.2 mm, shortly ciliolate at margin, ciliolate at base. |
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42 S. laxistrobila |
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44 (42) |
Leaves denticulate at margin. |
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43 S. tama-montana |
+ |
Leaves long ciliolate at margin |
|
(45) |
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45 (44) |
Strobili less than 4 mm; sporophylls plane. |
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44 S. prostrata |
+ |
Strobili usually over 8 mm, up to 25 mm; larger ventral sporophylls navicular. |
|
45 S. longistrobilina |
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46 (41) |
Main stems erect or suberect; rhizophores restricted to basal or lower portion of main stems |
|
(47) |
+ |
Main stems prostrate or at least lower part prostrate, branches creeping or erect; rhizophores at intermittent intervals |
|
(55) |
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47 (46) |
Plants more than 30 cm high |
|
(48) |
+ |
Plants usually less than 30 cm high |
|
(49) |
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48 (47) |
Plants erect, 40-75 cm high; rhizophores restricted to basal portion of main stems. |
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46 S. decipiens |
+ |
Plants suberect, 30-40 cm high; rhizophores present in lower part of main stems. |
|
47 S. bodinieri |
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49 (47) |
Sporophylls strongly dimorphic, ventral sporophylls up to 1/2 as long as dorsal ones |
|
(50) |
+ |
Sporophylls not strongly dimorphic, ventral sporophylls ca. 2/3 as long as dorsal ones |
|
(52) |
|
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50 (49) |
Plants up to 30 cm high; main stems robust; dorsal leaves obovate-aristate. |
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48 S. pennata |
+ |
Plants 5-15(-25) cm high; main stems thin |
|
(51) |
|
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51 (50) |
Ventral leaves up to 3 mm; sporophylls minutely denticulate. |
|
49 S. leptophylla |
+ |
Ventral leaves up to 3 mm; sporophylls sparsely ciliate. |
|
50 S. minutifolia |
|
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52 (49) |
Main stems tuberous at base. |
|
51 S. chrysocaulos |
+ |
Main stems not tuberous at base |
|
(53) |
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53 (52) |
Axillary leaves ovate-deltoid. |
|
52 S. effusa |
+ |
Axillary leaves ovate-lanceolate |
|
(54) |
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54 (53) |
Plants erect, main stems simple, branched in upper portion; leaves on stems and branches rather distant; rhizophores restricted to basal portion of main stems; base of ventral leaves shortly ciliolate. |
|
53 S. labordei |
+ |
Plants ascending to erect, main stems branched in lower and upper portions; leaves on main stems rather approximate; rhizophores on lower portion of main stems; base of ventral leaves long ciliolate. |
|
54 S. repanda |
|
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|
55 (46) |
Stems long creeping, fertile branches not erect, plants up to 30 cm or more; rhizophores on main stems and branches at intermittent intervals |
|
(56) |
+ |
Main stems and branches prostrate or fertile branches suberect, plants usually less than 20 cm, or main stems erect, normally less than 25 cm high; rhizophores on prostrate stems or restricted to basal portion of erect main stems |
|
(61) |
|
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56 (55) |
Dorsal leaves obovate, long ciliolate in upper portion. |
|
55 S. bisulcata |
+ |
Dorsal leaves not as above |
|
(57) |
|
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57 (56) |
Ultimate branches including leaves 10-15 mm wide. |
|
56 S. megaphylla |
+ |
Ultimate branches including leaves less than 8 mm wide |
|
(58) |
|
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58 (57) |
Dorsal leaves long aristate; sporophyll-pteryx of dorsal sporophylls not reaching apex (incomplete) |
|
(59) |
+ |
Dorsal leaves acuminate or shortly aristate; sporophyll-pteryx of dorsal sporophylls reaching apex or almost so (complete) |
|
(60) |
|
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59 (58) |
Basiscopic margin of ventral leaves entire. |
|
57 S. ornata |
+ |
Basiscopic margin of ventral leaves ciliolate. |
|
58 S. amblyphylla |
|
|
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|
60 (58) |
Leaves glabrous. |
|
59 S. monospora |
+ |
Leaves scabrous. |
|
60 S. trichophylla |
|
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|
61 (55) |
Prostrate stems very short, thin; plants small; fertile stems prominent |
|
(62) |
+ |
Plants not as above |
|
(63) |
|
|
|
|
62 (61) |
Sporangia only on ventral side of strobilus; sporophylls denticulate. |
|
61 S. devolii |
+ |
Sporangia on both sides of strobilus; sporophylls long ciliolate. |
|
62 S. ciliaris |
|
|
|
|
63 (61) |
Acroscopic base of ventral leaves not long ciliolate; leaves denticulate or shortly ciliolate at margin |
|
(64) |
+ |
Acroscopic base of ventral leaves long ciliolate |
|
(68) |
|
|
|
|
64 (63) |
Dorsal leaves rounded or obtuse at apex; leaves denticulate at margin. |
|
63 S. kouytcheensis |
+ |
Dorsal leaves acuminate or aristate; leaves denticulate or shortly ciliolate at margin |
|
(65) |
|
|
|
|
65 (64) |
Fertile branches short, complanate with sterile main stems and branches, creeping or ascending |
|
(66) |
+ |
Fertile branches long, erect |
|
(67) |
|
|
|
|
66 (65) |
Leaves shortly ciliolate at margin. |
|
64 S. boninensis |
+ |
Leaves minutely denticulate at margin. |
|
65 S. heterostachys |
|
|
|
|
67 (65) |
Dorsal leaves cordate at base, margin shortly ciliolate. |
|
53 S. labordei |
+ |
Dorsal leaves not cordate at base, margin minutely denticulate. |
|
65 S. heterostachys |
|
|
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|
68 (63) |
Fertile branches erect; ventral leaves involute in dry conditions |
|
(69) |
+ |
Fertile branches creeping; ventral leaves not involute in dry conditions |
|
(71) |
|
|
|
|
69 (68) |
Basiscopic base of ventral leaves with few long cilia. |
|
66 S. kurzii |
+ |
Basiscopic base of ventral leaves shortly ciliolate, elsewhere subentire |
|
(70) |
|
|
|
|
70 (69) |
Sporophylls obviously dimorphic. |
|
67 S. xipholepis |
+ |
Sporophylls slightly dimorphic. |
|
68 S. vaginata |
|
|
|
|
71 (68) |
Ventral leaves oblong-falcate; dorsal leaves ovate-lanceolate. |
|
69 S. drepanophylla |
+ |
Ventral leaves not oblong-falcate; dorsal leaves ovate or suborbicular |
|
(72) |
|
|
|
|
72 (71) |
Leaves not white-margined, sparsely ciliolate. |
|
70 S. chaetoloma |
+ |
Leaves distinctly white-margined |
|
(73) |
|
|
|
|
73 (72) |
Ventral leaves denticulate at margin, bearing several long hairs at basal portion. |
|
71 S. lutchuensis |
+ |
Ventral leaves long ciliolate at margin. |
|
72 S. albociliata |