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FOC | Family List | FOC Vol. 19 | Cucurbitaceae

11. Thladiantha Bunge, Enum. Pl. Chin. Bor. 29. 1833.

赤瓟属 chi bao shu

Authors: Anmin Lu & Charles Jeffrey

Herbs, perennial, climbing, scandent, or prostrate. Roots usually tuberous. Stems herbaceous, scabrous. Tendrils simple or 2-fid. Leaves mostly simple, vary rarely pedately 3-7-foliolate; leaf blade cordate, margin dentate. Plants dioecious. Male flowers in cymes-racemes or rarely solitary; calyx tube broadly campanulate or cupular; segments 5, linear, lanceolate, ovate-lanceolate, or oblong, 1-3-veined; corolla yellow, campanulate, 5-partite; segments oblong, broadly ovate, or obovate, usually 5-7-veined, entire; stamens 5, inserted on calyx tube, free, often 4 in pairs, other 1 free; filaments short; anthers oblong or ovoid, all 1-celled; anther cell erect; rudimentary ovary glandlike. Female flowers solitary or 2- or 3-fascicled on a short peduncle; calyx and corolla as in male flowers; ovary ovoid, oblong, or fusiform, smooth or verrucose; ovules numerous, horizontal; style 3-fid; stigmas 3, 2-lobed, reniform. Fruit fleshy, indehiscent, smooth or verrucose, many ribbed or not ribbed. Seeds numerous, horizontal.

Twenty-three species: Bhutan, China, India, Indonesia, Japan (introduced), Korea, Laos, Myanmar, Nepal, Thailand, Vietnam; 23 species (19 endemic) in China.

1 Bracts in male flowers flabelliform, dentate   (2)
+ Male flowers without flabelliform bracts   (4)
2 (1) Calyx segments entire, 0.6-3 mm wide.   1 T. cordifolia
+ Calyx segments dentate or irregularly lobed, (3-)4-11 mm wide   (3)
3 (2) Calyx segments (3-)4-6 mm wide, with 1-3 triangular lobes.   2 T. grandisepala
+ Calyx segments to 11 mm wide, flabelliform, margin irregularly lobed.   3 T. medogensis
4 (1) Ovary and fruit scaly or verrucose   (5)
+ Ovary and fruit not scaly or verrucose   (7)
5 (4) Leaf blade ovate-lanceolate or narrowly ovate-triangular; ovary and fruit verrucose.   23 T. longifolia
+ Leaf blade ovate-cordate; fruit with thick scales   (6)
6 (5) Ovary and fruit ovoid-globose, apex with a long rostrum, scales at base of fruit less than 5 mm, not adnate to pedicel.   21 T. pustulata
+ Ovary and fruit ovoid-oblong, apex without a rostrum, scales at base of fruit to 10 mm, adnate to pedicel.   22 T. henryi
7 (4) Leaves palmately deeply 5-lobed.   20 T. palmatipartita
+ Leaves not palmately deeply 5-lobed   (8)
8 (7) Male flowers congested at apex of rachis into pseudocapitulum   (9)
+ Male flowers in cyme-panicle, panicle, raceme, or rarely solitary, but not capitellate   (10)
9 (8) Calyx segments lanceolate, apex acute, 1-veined.   18 T. capitata
+ Calyx segments linear, apex obtuse, 3-veined.   19 T. davidii
10 (8) Plants sparsely hairy or subglabrous   (11)
+ Plants with dense multicellular setae, very rarely subglabrous   (15)
11 (10) Calyx segments 1-veined   (12)
+ Calyx segments 3-veined   (13)
12 (11) Basal lateral vein along base of leaf blade; calyx segments linear, over 9 mm.   13 T. oliveri
+ Basal lateral vein not along base of leaf blade; calyx segments triangular- lanceolate, 4-5 mm.   14 T. montana
13 (11) Tendrils 2-fid, leaves not lobed or lobed.   17 T. dentata
+ Tendrils simple, leaves not lobed   (14)
14 (13) Leaf blade broadly ovate-cordate; tendrils filiform; pedicel in male flowers 1.2-2.5 cm; fruit with a long beak.   15 T. maculata
+ Leaf blade narrowly ovate or narrowly ovate-lanceolate; tendrils robust; pedicel 0.5-1 cm; fruit without beak.   16 T. punctata
15 (10) Leaves simple, unlobed or margin lobed or serrate, or pedately 3-5-foliolate.   4 T. hookeri
+ Leaves unlobed, rarely irregularly lobed   (16)
16 (15) Tendrils simple   (17)
+ Tendrils 2-fid   (19)
17 (16) Male flowers solitary or in a short raceme, bracts leaflike.   5 T. dubia
+ Male flowers without bracts   (18)
18 (17) Petiole less than 1.5 cm, leaf blade lanceolate or narrowly ovate.   6 T. sessilifolia
+ Petiole over 2 cm, leaf blade ovate-cordate.   7 T. villosula
19 (16) Male flowers dimorphic, solitary, and in raceme with (2 or)3(-5) flowers in axil of same leaf.   8 T. dimorphantha
+ Male flowers in panicle or raceme   (20)
20 (19) Stem and leaves with pale yellow multicellular setae or very rarely subglabrous, sinus at base of leaf closed; fruit orange-red when mature.   9 T. nudiflora
+ Stem and leaves with gray-white multicellular setae, sinus at base of leaf open; fruit whitish yellow when mature   (21)
21 (20) Calyx segments broadly lanceolate or obovate-oblong, 10-15 mm.   10 T. longisepala
+ Calyx segments triangular-lanceolate, less than 10 mm   (22)
22 (21) Ovary and fruit with dense yellow-brown spiniform setae; seeds 6.5-7 mm, sparsely irregularly verrucose on surface.   11 T. setispina
+ Ovary and fruit with long setae; seeds 4.5-5 mm, not verrucose on surface.   12 T. lijiangensis

  • List of lower taxa


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