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FOC | Family List | FOC Vol. 11 | Zygophyllaceae

1. Zygophyllum Linnaeus, Sp. Pl. 1: 385. 1753.

霸王属 ba wang shu

Sarcozygium Bunge.

Shrubs, perennial herbs, or rarely annuals. Leaves opposite, even-pinnate or rarely simple; petiole herbaceous or membranous; leaflet blades plane or sticklike, fleshy. Flowers axillary, solitary or paired. Sepals 4 or 5, sometimes caducous. Petals as many as sepals, orangish red, white, or yellow, sometimes with an orange claw or pale along margin, rarely apetalous. Stamens 8-10, abaxially with scale-form appendages. Ovary 3-5-locular. Capsule winged or not, dehiscent or indehiscent, with 1 to several seeds per carpel.

More than 100 species: arid and semiarid regions of N Africa, C, N, and W Asia, Australia, S Europe; 19 species (two endemic) in China.


1 Fruit conspicuously winged   (2)
+ Fruit sometimes ridged but not conspicuously winged   (9)
       
2 (1) Shrubs; sepals and petals 4   (3)
+ Herbs; sepals and petals 5   (4)
       
3 (2) Capsule nearly spherical, 1.8-4 cm.   2 Z. xanthoxylon
+ Capsule narrowly ovoid to obovoid, 1-1.6 cm.   3 Z. kaschgaricum
       
4 (2) Leaflets 1 pair; capsule spherical, ca. 1 cm.   14 Z. fabagoides
+ Leaflets 1-5 pairs; capsule not spherical, or if spherical larger than 1 cm   (5)
       
5 (4) Capsule spherical, width of wings more than 5 mm   (6)
+ Capsule not spherical, width of wings less than 3 mm   (7)
       
6 (5) Leaves with 2 or 4 leaflets; leaflet blades 1-2.5 × 0.5-2 cm.   15 Z. potaninii
+ Leaves with 6-10 leaflets; leaflet blades 0.5-1.2 × 0.2-0.8 cm.   16 Z. macropterum
       
7 (5) Stamens ca. 2 × as long as petals; leaflet blades ovate to rotund.   17 Z. iliense
+ Stamens shorter than petals; leaflet blades linear, linear-oblong, or lanceolate   (8)
       
8 (7) Capsule lanceolate, base cuneate, apex acuminate.   18 Z. oxycarpum
+ Capsule oblong-ovoid to ovoid, both ends obtuse.   19 Z. pterocarpum
       
9 (1) Fruit not more than 2 × as long as wide, indehiscent.   1 Z. gobicum
+ Fruit more than 2 × as long as wide, dehiscent   (10)
       
10 (9) Leaves with more than 2 leaflets at least for majority of leaves   (11)
+ Leaves with only 2 leaflets   (13)
       
11 (10) Plants prostrate to ascending; leaflet blades 1-3 mm wide.   13 Z. mucronatum
+ Plants erect or spreading; leaflet blades 3-15 mm wide   (12)
       
12 (11) Perennial herbs; leaflet blades 3-5 mm wide; sepals 7-8 mm; capsule 2.5-4 cm, falcately curved.   11 Z. jaxarticum
+ Annual or biennial herbs; leaflet blades 4-15 mm wide; sepals 5-6 mm; capsule 1.6-2.5 cm, straight or slightly curved but not falcate.   12 Z. loczyi
       
13 (10) Stamens shorter than petals   (14)
+ Stamens longer than petals   (15)
       
14 (13) Plants 7-15 cm tall; petiole scabrous; pedicel 1-3 mm.   4 Z. kansuense
+ Plants 30-80 cm tall; petiole not scabrous; pedicel 10-18 mm.   5 Z. obliquum
       
15 (13) Plants 10-20 cm tall; capsule apex acuminate.   10 Z. rosowii
+ Plants 20-60(-100) cm tall; capsule apex not acuminate   (16)
       
16 (15) Capsule more than 6 mm in diam.; leaflet blades 1.5-3 cm wide   (17)
+ Capsule less than 5 mm in diam.; leaflet blades 0.7-1 cm wide   (18)
       
17 (16) Capsule cylindric, both ends not slender, 0.7-1 cm in diam.; seed surface smooth, margin entire; plants 40-80 cm tall.   6 Z. macropodum
+ Capsule oblanceolate to oblong, 1-1.5 cm in diam.; seed surface with pores, margin irregular; plants 20-25 cm tall.   7 Z. sinkiangense
       
18 (16) Leaflet blades oblong to oblanceolate; plants erect.   8 Z. brachypterum
+ Leaflet blades obovate to oblong-obovate; plants spreading or prostrate.   9 Z. fabago

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