15. Cyrtomium tukusicola Tagawa, Acta Phytotax. Geobot. 7(2): 79. 1938.
齿盖贯众 chi gai guan zhong
Cyrtomium macrophyllum (Makino) Tagawa var. tukusicola (Tagawa) Tagawa.
Plants 40-60 cm tall. Rhizome erect, densely covered with lanceolate blackish brown scales. Stipe stramineous, 18-28 cm, 2-3 mm in diam. at base, lower portion densely scaly; scales blackish brown, ovate or lanceolate, toothed, often twisted. Lamina oblong-ovate to oblong-lanceolate, 24-50 × 14-20 cm, base not contracted or slightly wider, 1-imparipinnate, apex obtuse; rachis with sparse lanceolate to linear brown scales. Lateral pinnae 2-8 pairs, alternate, slightly ascendant, shortly stalked, basal 1 or 2 pairs ovate and larger, remaining pinnae oblong-lanceolate or narrowly ovate, entire, sometimes acroscopic margins sparsely serrulate; middle pinnae 11-15 × 3-5 cm, base broadly cuneate or rounded-cuneate, apex acuminate or caudate; terminal pinna obovate or rhombic-ovate, forked or trifurcate, 7-14 × 4-10 cm; hard papery, glabrous on both surfaces; venation pinnate, slightly raised abaxially, indistinct adaxially, veinlets anastomosing to form multiple rows of areoles. Sori throughout abaxial surface of pinnae; indusia margins finely dentate.
Forests; 1000-2600 m. Chongqing (Nanchuan), Guizhou (Fanjing Shan, Kaili), Hunan (Longshan, Sangzhi), Sichuan (Cangxi, Emei, Leibo), C Taiwan, Yunnan (Yangbi), Zhejiang (Qingyuan) [Japan].
Cyrtomium tukusicola is an apomictic species and is sometimes treated as a variety of C. macrophyllum.