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6. Hylotelephium H. Ohba, Bot. Mag. (Tokyo). 90: 46. 1977.

八宝属 ba bao shu

Herbs perennial, glabrous or hairy. Roots fibrous or tuberous, often carrot-shaped. Rhizome short, fleshy or woody; young branches not covered with scales. Flowering stem arising from lateral bud of previous year, annual and abscissing from base or persisting and basally woody. Stem leaves alternate, opposite, or 3-5-verticillate; leaf blade flat, glabrous, spurless. Inflorescence terminal, sometimes also subterminal and thus paniculate, cymose, often corymbiform, sometimes umbel-like, many flowered, bracteate. Flowers bisexual or unisexual, unequally (4 or)5-merous, sometimes reduced. Sepals usually shorter than petals, base subconnate and spurless. Petals usually free, purple, red, pink, or white, occasionally yellowish or greenish. Stamens 2 × as many as petals, inserted basally on corolla. Nectar scales oblong-cuneate to linear-oblong, apex rounded or subemarginate. Carpels free, base narrow, substipitate. Follicles erect, many seeded. Seeds narrowly winged.

About 33 species: Asia, Europe, North America; 16 species (six endemic) in China.


1 Basal leaves 2   1 H. bonnafousii
+ Basal leaves not 2.   (2)
       
2 (1) Stems mostly ascending, less than 20 cm.   (3)
+ Stems erect, more than 30 cm.   (6)
       
3 (2) Leaves orbicular to orbicular-flabellate, 3-verticillate   5 H. sieboldii var. chinense
+ Leaves neither orbicular nor orbicular-flabellate, alternate, opposite, or 3-verticillate.   (4)
       
4 (3) Leaves alternate   2 H. tatarinowii
+ Leaves opposite or 3-verticillate.   (5)
       
5 (4) Leaves opposite, margin usually entire   3 H. ewersii
+ Leaves opposite or 3-verticillate, margin slightly crenulate   4 H. mongolicum
       
6 (2) Inflorescences both terminal and axillary, interrupted elongate in outline   6 H. angustum
+ Inflorescences all terminal or nearly so, corymbiform or capitate-umbellate in outline.   (7)
       
7 (6) Inflorescences capitate-umbellate   16 H. subcapitatum
+ Inflorescences corymbiform.   (8)
       
8 (7) Leaves usually 3-5-verticillate, sometimes basal ones opposite; leaf axil bulbils often present.   (9)
+ Leaves usually opposite or alternate, rarely 3-verticillate; leaf axil bulbils absent.   (10)
       
9 (8) Leaves usually longer than internodes, 4-8 × 2.5-3.5 cm   7 H. verticillatum
+ Leaves usually shorter than internodes, 2-4 × 0.7-1.2 cm   8 H. viviparum
       
10 (8) Leaves usually opposite, rarely 3-verticillate; anthers violet or purple.   (11)
+ Leaves usually alternate; anthers yellow (brown-red in H. tangchiense).   (13)
       
11 (10) Stamens not exserted from corolla   9 H. erythrostictum
+ Stamens exserted from corolla.   (12)
       
12 (11) Leaf base cuneate; petals purplish   10 H. spectabile
+ Leaf base subcordate and amplexicaul; petals reddish   11 H. pseudospectabile
       
13 (10) Stems ± flexuous; leaves elliptic-obovate or suborbicular.   (14)
+ Stems erect; leaves elliptic-lanceolate, oblong-ovate, or oblong.   (15)
       
14 (13) Basal stem leaves elliptic-obovate, to 8.5 × 3 cm, base attenuate; anthers yellow   12 H. mingjinianum
+ Basal stem leaves suborbicular, 5-10 × 4-9 cm, base abruptly narrowed; anthers brown-red   13 H. tangchiense
       
15 (13) Roots carrot-shaped; petals purplish red   15 H. triphyllum
+ Roots not carrot-shaped; petals white or reddish   14 H. pallescens

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