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FOC | Family List | FOC Vol. 20-21 | Asteraceae


斑鸠菊族 ban jiu ju zu

Authors: Yilin Chen & Michael G. Gilbert

Herbs, shrubs, sometimes climbing, or trees; hairs simple, T-shaped, or stellate. Leaves usually alternate [rarely opposite or whorled], leaf blade entire or serrate-dentate [rarely pinnately divided], venation pinnate, rarely with 3 basal veins (Distephanus). Synflorescences mostly terminal, less often terminal on short lateral branches or axillary, mostly cymose paniculate, less often spikelike, forming globose compound heads or reduced to a solitary capitulum. Capitula discoid, homogamous. Phyllaries generally imbricate, in several rows, rarely in 2 rows, herbaceous, scarious or leathery, outer gradually shorter. Receptacle flat or rather convex, naked or ± fimbriate. Florets 1-400, all bisexual, fertile; corolla tubular, purple, reddish purple, pink, or white, rarely yellow (Distephanus), limb narrowly campanulate or funnelform, 5-lobed. Anther base bifid, auriculate, acute or hastate, rarely caudate, apex appendaged. Style branches usually long and slender, apex subulate or acute, dorsally pilose, without appendage. Achenes cylindric or slightly flattened, (2-)5-10[-20]-ribbed, or 4- or 5-angled, rarely ± terete; pappus usually present, persistent, of many filiform setae, bristles, or scales, often 2-seriate with inner series of setae or bristles and shorter outer series of scales, sometimes very few and deciduous (Camchaya) or absent (Ethulia).

Up to 120 genera and 1,400 species: throughout the tropics and extending into some temperate regions; six genera (one introduced) and 39 species (ten endemic, two introduced) in China.

The classification of this tribe is still in flux with some authors not accepting most of the recent segregates from Vernonia. See Robinson and Skvarla (Taiwania 55: 254-272. 2010) for a treatment of the Chinese species of Vernonieae using these segregates.

1 Capitula densely clustered into compound synflorescences subtended by (1-)3 leaflike bracts, each involucre with 1-4 florets and ca. 8 phyllaries   (2)
+ Capitula in lax panicles, rarely in sessile clusters and then each involucre with more than 4 florets and always with many phyllaries   (3)
2 (1) Capitula (1-)10-40 per cluster, clusters terminal, solitary or in flat-topped synflorescences, each subtended by (2 or)3 ± triangular bracts; pappus with many straight, scabrid-barbellate, basally widened awns.   90 Elephantopus
+ Capitula 1-5+ per cluster, clusters axillary in long spikelike synflorescences, each subtended by 1 or 2 lanceolate to spatulate or linear bracts; pappus with 2 very long, bent or curved awns.   91 Pseudelephantopus
3 (1) Corolla golden yellow; leaves basally 3-veined.   88 Distephanus
+ Corolla pale to dark purple, pink, or white; leaves pinnately veined   (4)
4 (3) Pappus absent; achenes (2-)4- or 5(or 6)-ribbed, apex truncate or concave with an enlarged rim.   86 Ethulia
+ Pappus present [or absent]; achenes 10-ribbed, 4- or 5-angled, or terete, apex rounded   (5)
5 (4) Pappus of 1-10 easily deciduous bristles [or absent].   89 Camchaya
+ Pappus long, copious, persistent, outermost setae sometimes short and scalelike.   87 Vernonia

Lower Taxa


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