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FOC | Family List | FOC Vol. 20-21 | Asteraceae

16. Himalaiella Raab-Straube, Willdenowia. 33: 390. 2003.

须弥菊属 xu mi ju shu

Authors: Zhu Shi & Eckhard von Raab-Straube

Saussurea sect. Elatae J. D. Hooker, Fl. Brit. India 3: 373. 1881.

Herbs, biennial or perennial, tall or rarely stemless. Stems usually densely leafy or rarely scapiform. Basal leaves often lyrately pinnately lobed to pinnately divided. Capitulum solitary and terminal on stem or in a racemiform, paniculiform, or corymbiform synflorescence, often nodding or sometimes erect, sessile or pedunculate. Involucre hemispheric to campanulate. Phyllaries sometimes reflexed. Receptacle densely covered with subulate alveolate bristles and rarely also with subulate paleae. Corolla reddish purple, purple, pink, or white, gland-dotted. Anther tails moderately lacerate. Achene blackish, 2-5.5 mm, 4- or 5-angled, glabrous, wrinkled, squamulate, or muricate, apical rim forming a conspicuous crown. Pappus homomorphic, in 1 row; bristles snow white, dirty white, or brownish, plumose, basally connate into a ring, caducous as a whole.

Thirteen species: Afghanistan, Bhutan, China, N India, Kashmir, Laos, Myanmar, Nepal, Pakistan, N Thailand, N Vietnam; SW Asia (Iran); seven species (one endemic) in China.

In C. Shih and S. Y. Jin (FRPS 78(2): 57-66. 1999), most species of Himalaiella are included in Saussurea subg. Frolovia (Candolle) Lipschitz. They are, however, more closely related to Jurinea than to Saussurea.

1 Leaf blade of basal and lower stem leaves pinnately or bipinnately divided, terminal lobe not much larger than lateral ones   (2)
+ Leaf blade of basal and lower stem leaves lyrately pinnately lobed or lyrately pinnately divided, terminal lobe much larger than lateral ones or blade undivided   (3)
2 (1) Plants almost stemless; capitula few, clustered in center of leaf rosette.   1 H. yakla
+ Plants with tall stems; capitula many, in a racemiform or paniculiform synflorescence.   7 H. peguensis
3 (1) Stem simple, scapiform with reduced leaves; capitulum always solitary; leaf blade adaxially grayish green, arachnoid tomentose.   2 H. abnormis
+ Stem branched or rarely simple, leafy, not scapiform; capitula numerous or solitary; leaf blade adaxially green, scabrous, strigose, or glabrescent   (4)
4 (3) Receptacle with bristles and long subulate paleae; pappus brown.   3 H. auriculata
+ Receptacle with bristles but without paleae; pappus white or straw-colored   (5)
5 (4) Plants few branched, with 2-4 capitula; outer phyllaries without appendage, long acuminate.   4 H. qinghaiensis
+ Plants usually much branched, capitula numerous; outer phyllaries with a green, herbaceous appendage   (6)
6 (5) Involucre 1-2 cm in diam.; capitula solitary or clustered, subsessile or shortly pedunculate; corolla 1-1.2 cm; pappus 0.8-1 cm.   6 H. nivea
+ Involucre 2-4 cm in diam.; capitulum mostly solitary, long pedunculate; corolla 1.5-2 cm; pappus 1.4-1.6 cm.   5 H. deltoidea

Lower Taxa


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