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Annual or biennial herbs; main root solitary, vertical, tapered or ± fusiform; basal leaves (including petiole) usually less than 8 cm |
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(2) |
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Perennial herbs, subshrubs, or shrubs; roots several, not fusiform (if biennials with single fusiform root then basal leaves 11-18 cm); basal leaves usually more than 8 cm |
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(7) |
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2 (1) |
Synflorescence a simple erect terminal raceme; capitula 5-10(-15) mm in diam. |
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1 A. macrocephala |
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Synflorescence a panicle with spreading lateral branches; capitula 1.5-4(-6) mm in diam |
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(3) |
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3 (2) |
Capitula 3-6 mm in diam.; marginal female florets 5-30, bisexual florets 30-120 |
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(4) |
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Capitula 1.5-3(-4) mm in diam.; marginal female florets 3-6, bisexual florets 18-28 |
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(5) |
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4 (3) |
Plants 50-150 cm tall; middle stem leaf blades 4-8(-13) × 3-6(-15) cm; female florets 20-30, often in double series, bisexual florets 80-120. |
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2 A. sieversiana |
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Plants 25-40 cm tall; middle stem leaf blades 1.5-2 × 1-1.5 cm; female florets 5-20 in single series, bisexual florets 30-50. |
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22 A. succulentoides |
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5 (3) |
Basal leaf blades 8-15 × 8-15 cm. |
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19 A. nakaii |
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Basal leaf blades 3-4.5(-5) × 2-4 cm |
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(6) |
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6 (5) |
Capitula 2-3(-4) mm in diam.; bisexual florets 18-28. |
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20 A. anethifolia |
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Capitula 1.5-2(-2.5) mm in diam.; bisexual florets 8-16. |
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21 A. anethoides |
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7 (1) |
Disk florets 8-20 |
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(8) |
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Disk florets 20-80 |
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(11) |
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8 (7) |
Plants 15-25(-30) cm tall, densely gray or yellowish tomentose. |
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16 A. younghusbandii |
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Plants 25-80 cm tall, gray or yellowish arachnoid tomentose or silky pubescent, lower parts sometimes glabrescent |
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(9) |
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9 (8) |
Middle stem leaf blades 3-5 × 3-5 cm; marginal female florets 2-5. |
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18 A. fauriei |
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Middle stem leaf blades 0.5-2 × 0.8-2.8 cm; marginal female florets 4-10 |
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(10) |
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10 (9) |
Ultimate leaf lobules narrowly spatulate, linear, or linear-oblanceolate, 1-3 × 0.5-1.5 mm. |
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9 A. xerophytica |
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Ultimate leaf lobules elliptic-oblanceolate or elliptic-lanceolate, 6-12 × 1-1.5 mm. |
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10 A. rutifolia |
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11 (7) |
Leaf blade entire or with apex crenately 3-5-lobed. |
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12 A. lagocephala |
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Leaf blade 1-3-pinnatisect |
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(12) |
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12 (11) |
Capitula 2-4(-5) mm in diam |
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(13) |
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Capitula 4-10 mm in diam |
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(16) |
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13 (12) |
Basal to middle stem leaves with petiole 2-12 cm; plants 60-150 cm tall. |
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5 A. absinthium |
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Basal to middle stem leaves sessile or with petiole to 2 cm; plants 20-70 cm tall |
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(14) |
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14 (13) |
Middle stem leaves sessile, gland-dotted. |
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11 A. obtusiloba |
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Middle stem leaves with petiole 5-20 mm, not gland-dotted |
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(15) |
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15 (14) |
Bisexual florets 20-30; perennial herbs or occasionally subshrubs with slender rootstock; plants densely yellowish or pale yellow sericeous. |
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6 A. frigida |
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Bisexual florets 35-50; subshrubs with a thick woody rhizome ca. 1 cm or more in diam.; plants arachnoid puberulent, glabrescent. |
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23 A. persica |
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16 (12) |
Plants densely yellowish to yellow sericeous-tomentose; corolla with multicellular hairs; basal leaves 5-10 cm, with 7-13 pairs of lateral segments. |
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17 A. stracheyi |
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Plants pubescent or glabrescent; corolla glabrous; basal leaves less than 5 cm, with 3-7 pairs of lateral segments, or leaves 3-cleft |
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(17) |
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17 (16) |
Plants 10-20 cm tall |
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(18) |
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Plants 20-70 cm tall |
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(20) |
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18 (17) |
Middle stem leaves with petiole 2-3 cm, blade pubescent, later glabrescent, greenish, abaxially gland-dotted; phyllary margin black scarious. |
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13 A. disjuncta |
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Middle stem leaves with petiole 0.4-1 cm, blade densely gray or yellowish tomentose, or glabrescent, abaxially not gland-dotted; phyllary margin brown scarious |
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(19) |
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19 (18) |
Perennial herbs; middle stem leaves with (3 or)4(or 5) pairs of segments, segments serrate or with lobules 2-8 × 0.6-1 mm, pubescent, soon glabrescent; phyllaries with brown scarious margin, glabrescent. |
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14 A. niitakayamensis |
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Caespitose dwarf subshrubs; middle stem leaves with 2(or 3) pairs of segments, 3-5-lobulate, lobules 1-2 × 0.5-1 mm; leaves and phyllaries densely yellowish or gray tomentose. |
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15 A. minor |
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20 (17) |
Middle stem leaves (1-)1.5-4 cm, densely yellowish or white sericeous-tomentulose; middle lobes 1.5-4 mm wide |
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(21) |
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Middle stem leaves 0.8-1.5 cm, tomentose, pubescent, or glabrescent; middle lobe less than 1 mm wide |
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(22) |
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21 (20) |
Plants 40-60(-75) cm tall; middle stem leaves elliptic or ovate; phyllary with narrow, white scarious margin. |
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3 A. sericea |
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Plants 20-40(-50) cm tall; middle stem leaves suborbicular or orbicular-reniform; phyllary with wide, brown scarious margin. |
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4 A. aschurbajewii |
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22 (20) |
Middle stem leaves with 5-7 pairs of segments, distal lobes pinnatisect or 3-lobulate, proximally not lobed, pubescent, glabrescent; phyllaries pubescent. |
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8 A. rupestris |
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Middle stem leaves with 2 or 3(-5) pairs of segments, segments parted or lobulate; leaves and phyllaries yellowish sericeous-tomentose or pubescent or glabrescent. |
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7 A. argyrophylla |