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Moss China | Family List | Moss China V. 1 | Dicranaceae | Dicranodontium

8. Dicranodontium uncinatum (Harv.) Jaeg., Ber. Thätigk. St. Gallischen Naturwiss. Ges. 1877–78: 380. 1880; Thysanomitrion uncinatum Harv. in Hook., Icon. Pl. 1: 22. 1836.


Dicranum uncinatum (Harv.) C. Müll., Syn. Musc. Frond. 1: 404. 1848, hom. illeg. Dicranodontium blindioides (Besch.) Broth., Nat. Pflanzenfam. 1(3): 336. 1901. Dicranum blindioides Besch., Rev. Bryol. 18: 88. 1891. Type. China: Yunnan, Lopin-chan (Lan-Kong), Delavay 4815 (holotype H). Synonymized by Frahm (1997). Dicranodontium blindioides var. robustum Broth. in Handel Mazzetti, Symb. Sin. 4: 20. 1929. syn. nov. Type. China: Yunnan, 27o 53´N, alt. 2130 m, Handel-Mazzetti 9333 (holotype H). Dicranodontium circinatum (Milde) Schimp., Syn. Musc. (ed. 2): 100. 1876. Dicranum longirostre var. circinatum Milde, Bot. Zeit 28: 417. 1870. Dicranodontium fleischerianum Schultze-Motel, Taxon 12: 127. 1963. Dicranodontium fleischerianum var. clemensiae (Bartr.) Schultze-Motel, Taxon 12: 127. 1963. Dicranodontium nitidum var. clemensiae Bartr., Philippine J. Sci. 61: 236. 1936. Dicranodontium nitidum (Dozy & Molk.) Fleisch., Musci Buitenzorg 1: 87. 1904, hom. illeg. Campylopus nitidus Dozy & Molk., Musci Frond. Ined. Archip. Ind. 5: 139. 1847.

Plants coarse, fairly robust, up to 7(–10) cm high, yellowish green to pale brownish, somewhat silky, in loose or dense tufts. Stems erect to flexuose or curved, simple or sparsely branched; central strand present. Leaves up to 8(–12) mm long, strongly falcate-secund at upper stems, scarcely altered when dry, lanceolate, rather abruptly narrowed from an oblong-ovate, sheathing base to a long, subulate to semi-canaliculate acumen, sharply pointed at the apex; margins somewhat involute, entire below, serrulate near the apex; costa yellowish to reddish brown, occupying ¼ – 1/3 the leaf base width, excurrent in a short awn, slightly longitudinally furrowed and serrulate at back above; upper cells linear, thick-walled, distinctly projecting at the upper ends; lower cells rectangular, 38–70 µm × 15–20 µm, thick-walled and porose, pale yellowish, forming 5–7 rows of linear cells at the margins; alar cells inflated, thin-walled, forming well marked auricles, sometimes fragile, hyaline or reddish brown. Dioicous. Setae 8–12 mm long, yellowish, strongly flexuose when dry, cygneous when moist; capsules cylindric, brownish, ca. 2 mm × 0.7 mm, exothecial cells rectangular, slightly thick-walled; peristome teeth divided nearly to the base, reddish brown, vertically striate below; papillose at the tips. Spores 16–17 µm in diameter, pale yellowish.

Chinese specimens examined: ANHUI: She-xian Co., W.-F. Zhen 1124 (IFSBH). FUJIAN: Wuyi Shan (Mt.), D.-K. Li 10980 (SHM). GUANGDONG: Ru-yuan Co., Yue 73 Collecting Team 417 (IBSC, IFSBH). GUANGXI: Long-sheng Co., P.-C. Wu 654(b) (PE); Xing-an Co., C. Gao & G.-C. Zhang 1404, 1600 (IFSBH). HAINAN: Diao-luo Shan (Mt.), Yue 77 Collecting Team 2919 (IBSC, IFSBH); Ding-an Co., Z. Wang 35628 (IBSC, IFSBH). HUNAN: Hengyang City, T. Cao 22065 (ALBT, IFSBH). JIANGXI: Lu Shan (Mt.), L.-J. Bi 20 (IFSBH). SICHUAN: Da-jin Chuan Co., Z.-T. Guan 5330 (MO, as Dicranodontium circinatum); Lei-bo Co., Z.-T. Guan 8810 (MO, as Dicranodontium circinatum); Li-xian Co., Z. Wang s.n. (IFSBH); Mu-li Co., C. Gao et al. 20726, 21073 (IFSBH); Tian-quan Co., C. Gao et al. 18343 (IFSBH, MO). TAIWAN: Ilan Co., C.-C. Chuang 2060 (MO). XIZANG: Mi-lin Co., S.-K. Chen 168(a) (IFSBH, KUN); Mo-tuo Co., Y.-G. Su 4472(a) (IFSBH, KUN); Ya-dong Co., M. Zang 101 (IFSBH, KUN). YUNNAN: Gong-shan Co., M.-Z. Wang 10590(c) (IFSBH, PE); Jing-dong Co., J.-K. Zhang 1242 (IFSBH, YUKU); Li-jiang Co., W.-X. Xu 716 (IFSBH, YUKU). ZHEJIANG: Nanyindan (Mt.), C. Gao 879 (ALBT, IFSBH); Sui-chang Co., Z.-L. Liu 613, 1171 (IFSBH, HSNU).

Habitat: on soil, rocks, grasslands, or rotten wood and bases of trees; Distribution: China, Bhutan, Nepal, India, Sri Lanka, Myanmar, Thailand, Vietnam, Indonesia, Malaysia, the Philippines, Japan, Europe, and North America.

Type of Dicranodontium blindioides var. robustum matches the concept of D. uncinatum very well and its distribution range falls in line with that of D. blindioides, which is also a synonym of the latter species.

Illustrations: C. Gao (ed.) 1994 (Pl. 82, figs. 8–15).


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