Plants green, yellowish green to brownish. Leafy stems simple, but usually with innovations from the axils of upper leaves, 8–18 mm long, 2.3–4.0 mm wide; axillary hyaline nodules well developed; central strand only slightly differentiated. Leaves in 18–38 pairs, densely arranged; middle to upper leaves linear-lanceolate to lanceolate, 2.0–2.7 mm × 0.30–45 mm, acuminate at apex, the upper half of leaves usually more or less rugose; base of dorsal laminae often rounded; vaginant laminae ca. 1/2 the leaf length, upper part equal to somewhat unequal; costa stout, slightly excurrent; margins crenulate; margins of apical and dorsal laminae forming a thick band 2–3 cells wide and 2–3 cells thick; margins of vaginant laminae forming a thinner band 2–3 cells wide and 1 cell thick; cells of apical and dorsal laminae subisodiametric, 7–9 µm wide, thick-walled, mammillose; cells of vaginant laminae similar to those of apical and dorsal laminae, but slightly larger and well demarcated with thicker cell walls. Sporophytes not seen.

Type. Indonesia: Java, Teysmann s.n.

Chinese specimens examined: GUANGDONG: Ding-hu Shan (Mt.), Z.-H. Li & P.-J. Lin 069 (IBSC, SYS). HAINAN: Diao-luo-shan (Mt.), Z.-H. Li 3122 (H, SYS); Jian-feng-ling (Mt.), P.-C. Chen et al. 481, 777 (PE, SYS); Ling-shui Co., Redfearn et al. 36101 (MO). TAIWAN: Nan-tou Co., S.-H. Lin 8506 (NICH, TUNG); Tai-bei Co., Iwatsuki 3671 (NICH). XIZANG: Mo-tuo Co., Y.-G. Su 3228 (HKAS, NICH). YUNNAN: W.-X. Xu 11280 (IBSC, SYS); Gong-shan Co., M.-Z. Wang 10604b, 10696b (PE, SYS).

Habitat: on moist soil and rocks; Distribution: China, Japan, Nepal, India, Sri Lanka, Myanmar, Thailand, Malaysia, Indonesia, the Philippines, and New Guinea.

Illustrations: Z.-H. Li 1985 (Fig. 17, m–v).