1 |
Leaves without costa |
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25 Fissidens hyalinus |
+ |
Leaves with costa |
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(2) |
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2 (1) |
Plants minute, budlike with very short stems |
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(3) |
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Plants larger, with distinct elongate stems |
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(4) |
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3 (2) |
Protonema persistent; costa very weak, lacking on lower leaves; cells of apical laminae 23-35(-45) µm long; calyptra smooth, |
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41 Fissidens protonematicola |
+ |
Protonema not persistent; costa distinct; cells of apical laminae 13-17(-21) µm long; calyptra distinctly scabrous by papillae at upper end of each surface cell |
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8 subsp. kiusiuensis |
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4 (2) |
Leaves soft; cells of apical laminae lax and large, 19-50 µm long |
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(5) |
+ |
Leaves more or less firm; cells of apical laminae dense, usually not more than 20 µm long, or if longer, the upper end of vaginant laminae extremely unequal |
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(6) |
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5 (4) |
Leaves densely arranged to the base of stems, without limbidia or weakly bordered by 1-2 rows of elongate cells (often submarginal) |
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4 Fissidens bogoriensis |
+ |
Leaves laxly arranged, at least not densely arranged to the base of stems, with distinct limbidia all around the margins |
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16 Fissidens flaccidus |
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6 (4) |
Leaves at least partially limbate |
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(7) |
+ |
Leaves not limbate |
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(29) |
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7 (6) |
Limbidia all around the leaf margins |
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(8) |
+ |
Limbidia confined to the vaginant laminae or present only on vaginant laminae of perichaetial leaves |
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(16) |
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8 (7) |
Capsules curved, asymmetrical |
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(9) |
+ |
Capsules straight to nearly straight, symmetrical |
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(10) |
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9 (8) |
Leaves lanceolate to narrowly lanceolate; upper end of vaginant laminae open to costa (extremely unequal); cells of apical laminae 17-35 µm long |
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3 Fissidens beckettii |
+ |
Leaves oblong-lanceolate to ovate-lanceolate; upper end of vaginant laminae equal to slightly unequal; cells of apical laminae 7-14 µm long |
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50 Fissidens tosaensis |
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10 (8) |
Laminal cells with a small papilla in center |
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1 Fissidens angustifolius |
+ |
Laminal cells smooth |
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(11) |
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11 (10) |
Cells at base of vaginant laminae up to 42 µm long, much longer than those of apical and dorsal laminae (9-21 µm long); axillary hyaline nodules well differentiated |
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(12) |
+ |
Cells at base of vaginant laminae up to 21 µm long, only slightly longer than those of apical and dorsal laminae (5-14 µm long); axillary hyaline nodules not or weakly differentiated |
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(13) |
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12 (11) |
Leaves lanceolate, synoicous |
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52 Fissidens zollingeri |
+ |
Leaves narrowly lanceolate, rhizoautoicous |
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46 Fissidens subbryoides |
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13 (11) |
Sterile and fertile stems often differentiated; inflorescences rhizoautoicous; perichaetial leaves much larger than lower leaves |
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12 Fissidens curvatus |
+ |
Sterile and fertile stems undifferentiated; inflorescences various; perichaetial leaves not well differentiated from lower stem leaves |
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(14) |
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14 (13) |
Limbidia weak to relatively distinct, usually colorless; base of dorsal laminae not decurrent |
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5 Fissidens bryoides |
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Limbidia usually distinct on apical laminae, yellowish on older leaves; base of dorsal laminae more or less decurrent, or not decurrent |
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(15) |
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15 (14) |
Plants 5-15 mm high; leaves 1.8-2.5 mm long; setae 2.5-5.0(-10.0) mm long, lateral or terminal; antheridia 120-170 µm long |
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32 Fissidens longisetus |
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Plants smaller, not more than 10 mm high; leaves 1.4-1.6 mm long; setae 3.5-4.0 mm long; antheridia (190-) 210-340 µm long |
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20 Fissidens geppii |
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16 (7) |
Leaf cells and exothecial cells distinctly unipapilllose |
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6 Fissidens capitulatus |
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Leaf cells variously papillose or smooth, exothecial cells smooth |
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(17) |
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17 (16) |
Laminal cells smooth |
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(18) |
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Laminal cells papillose |
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(20) |
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18 (17) |
Setae only 0.4-0.8 mm long; capsules usually immersed in upper leaves, urns 0.7-0.8 mm long; dioicous or rhizoautoicous |
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47 Fissidens subsessilis |
+ |
Setae more than 2.0 mm long; capsules well exserted above the upper leaves, urns 0.4-0.65 mm long; autoicous |
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(19) |
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19 (18) |
Central strand not differentiated; upper leaves oblong-ovate, 1.2-1.5 mm long, acute or obtusely acute at apex; costa ending below the apex; vaginant laminae 2/3 -4/5 the leaf length |
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13 Fissidens diversifolius |
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Central strand weakly differentiated; upper leaves lanceolate to narrowly lanceolate, 1.75-2.30 mm long, narrowly acute at apex; costa percurrent to shortly excurrent; vaginant laminae 1/2 - 2/3 the leaf length |
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42 Fissidens schusteri |
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20 (17) |
Leaf cells with 1 (rarely 2) small papillae |
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(21) |
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Leaf cells pluripapillose |
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(22) |
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21 (20) |
Upper leaves oblong lanceolate, acute at apex; cells of apical laminae quadrate to hexagonal, 9-21 µm long, slightly mammillose with 1 (rarely 2) small, indistinct papillae, thin-walled |
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43 Fissidens schwabei |
+ |
Upper leaves lanceolate, narrowly acute at apex; cells of apical laminae irregularly rounded hexagonal, 3.5-9.0 µm long, highly mammillose with 1 (rarely 2) distinct papillae, moderately thick-walled |
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10 Fissidens crenulatus |
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22 (20) |
Setae more or less scabrous; perigonia in axils of leaves |
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24 Fissidens hollianus |
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Setae always smooth; perigonia terminal on short branches at the base of female stems (if perigonia in axils of leaves, limbidia submarginal) |
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(23) |
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23 (22) |
Leaves densely overlapping to base; upper leaves narrowly lanceolate or linear-lanceolate |
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(24) |
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Leaves laxly arranged near base; upper leaves oblong-lanceolate to lanceolate |
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(26) |
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24 (23) |
Axillary hyaline nodules indistinct; limbidia weak, found only on lower half of vaginant laminae of upper and perichaetial leaves |
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31 var. obscurirete |
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Axillary hyaline nodules well or weakly differentiated; limbidia found on vaginant laminae of almost every leaf |
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(25) |
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25 (24) |
Axillary hyaline nodules well differentiated; rhizoautoicous; fertile and sterile stems differentiated and connected at base; setae terminal on short female stems |
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30 Fissidens kinabaluensis |
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Axillary hyaline nodules weakly differentiated; autoicous; fertile and sterile stems not differentiated; setae terminal on stems |
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51 Fissidens wichurae |
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26 (23) |
Leaf cells highly mammillose, with a cluster of 2-5 high papillae at center |
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26 Fissidens incognitus |
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Leaf cells not visibly mammillose, with papillae along the cell walls |
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(27) |
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27 (26) |
Leaf apices acute; costa percurrent to shortly excurrent |
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7 Fissidens ceylonensis |
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Leaf apices rounded to obtuse; costa ending below leaf apex |
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(28) |
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28 (27) |
Vaginant laminae unequal; the minor laminae narrowed distally, ending on or very close to the costa; peristome teeth erect when wet, undivided or incompletely divided |
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18 Fissidens gardneri |
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Vaginant laminae more or less equal; the minor laminae acute, ending on or very close to the leaf margins; peristome teeth inflexed when wet, divided more or less 2/3 their length |
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33 Fissidens minutus |
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29 (6) |
Several rows of cells at margins of apical laminae lighter in color and smooth, markedly differentiated from inner cells as a paler band |
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(30) |
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Marginal cells not as above, usually not differentiated from inner cells, or if differentiated, then only 1-2 rows of cells, or darker in color due to multi-stratose cells |
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(31) |
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30 (29) |
Leaves lanceolate; costa percurrent, paler margins 3-4 cells wide; cross section of apical laminae 1-2 cells thick; setae 5-8 mm long |
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14 Fissidens dubius |
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Leaves narrowly lanceolate; costs excurrent, paler margins 1-3 cells wide; cross section of apical laminae 1 cell thick; setae not more than 2 mm long |
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2 Fissidens anomalus |
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31 (29) |
Leaf margins dark in color, 2- 4 cells thick |
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(32) |
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Leaf margins not differentiated from other laminal cells |
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(34) |
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32 (31) |
Axillary hyaline nodules well differentiated; base of dorsal laminae rounded |
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28 Fissidens javanicus |
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Axillary hyaline nodules not differentiated; base of dorsal laminae wedge-shaped |
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(33) |
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33 (32) |
Plants smaller, stems 8-11 mm long; leaf apices crenulate or nearly entire; leaf margins 2 cells thick; base of dorsal laminae not decurrent |
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29 Fissidens jungermannioides |
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Plants larger, stems 18-60 mm long; leaf apices irregularly dentate; leaf margin 2-4 cells thick; base of dorsal laminae decurrent |
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34 Fissidens nobilis |
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34 (31) |
Plants aquatic; leaf bases distinctly decurrent on stems |
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(35) |
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Plants not aquatic; leaf bases not distinctly decurrent on stems |
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(37) |
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35 (34) |
Leaves more or less crispate when dry; apical laminae 1 cell thick; costa clearly defined; with 1 row of irregularly quadrate to rectangular, smooth, more or less pellucid cells along both sides of costa |
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19 Fissidens geminiflorus |
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Leaves almost straight when dry; apical laminae 1-6 cells thick; costa obscure; cells along both sides of costa not different from other laminal cells |
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(36) |
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36 (35) |
Leaves translucent under microscope; apical laminae 1-4 cells thick; two rows of rectangular guide cells visible through surface cells from middle to upper portion of costa |
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39 Fissidens perdecurrens |
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Leaves not translucent under microscope; apical laminae 1-6 cells thick; two rows of guide cells not visible as above |
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21 Fissidens grandifrons |
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37 (34) |
Plants very small, leafy stems only 0.8-1.7 mm long; margins of vaginant laminae more or less distinctly serrate |
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(38) |
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Plants small to larger, leafy stems usually more than 3 mm long; margins of vaginant laminae almost entire to serrulate or crenulate |
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(39) |
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38 (37) |
Margins of vaginant laminae evenly serrate; cells of apical laminae smooth |
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15 Fissidens flabellulus |
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Margins of vaginant laminae irregularly and sharply serrate; cells of apical laminae distinctly papillose |
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44 Fissidens serratus |
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39 (37) |
Axillary hyaline nodules very prominent |
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(40) |
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Axillary hyaline nodules not well differentiated |
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(42) |
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40 (39) |
Cells of apical laminae multipapillose |
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45 Fissidens subangustus |
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Cells of apical laminae mammillose |
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(41) |
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41 (40) |
Central strand not differentiated; leaf apices widely acute; costa percurrent or ending a few cells below apex |
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11a var. crispulus |
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Central strand weakly differentiated; leaf apices narrowly acute; costa shortly excurrent |
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11b var. robinsonii |
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42 (39) |
Cells of vaginant laminae with 3-4 papillae at corners |
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49 Fissidens teysmannianus |
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Cells of vaginant laminae not papillose at corners |
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(43) |
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43 (42) |
Cells of apical laminae smooth or slightly mammillose |
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(44) |
+ |
Cells of apical laminae papillose or distinctly mammillose |
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(49) |
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44 (43) |
Plants small; leafy stems 1.4-9.0 mm long |
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(45) |
+ |
Plants medium-sized to large; leafy stems 18.0-72.0 mm long |
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(48) |
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45 (44) |
Costa ending 6-9 cells below the leaf apex |
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22 Fissidens guangdongensis |
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Costa percurrent to shortly excurrent |
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(46) |
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46 (45) |
Costa usually curved above the vaginant laminae; the dorsal laminae slightly decurrent at the base; leaf cells rather thin-walled |
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17 Fissidens ganguleei |
+ |
Costa more or less straight above the vaginant laminae; the dorsal laminae not decurrent; leaf cells more or less thick-walled |
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(47) |
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47 (46) |
Leaves lanceolate; cells of apical laminae moderately thick-walled, 10-24 µm long; synoicous (polyoicous) |
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38 Fissidens pellucidus |
+ |
Leaves lanceolate to narrowly lanceolate; cells of apical laminae thick- to very thick-walled, lumen 8-13 µm long; dioicous |
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9 Fissidens crassinervis |
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48 (44) |
Leaf apex mostly mucronate, rarely obtusely acute; cells of apical laminae 11-21 µm long, walls clear |
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40 Fissidens polypodioides |
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Leaf apex obtusely acute; cells of apical laminae 10-13 µm long, walls obscure |
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36 Fissidens obscurus |
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49 (43) |
Setae lateral or basal; perichaetial leaves much smaller than stem leaves |
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48 Fissidens taxifolius |
+ |
Setae terminal; perichaetial leaves not or indistinctly differentiated |
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(50) |
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50 (49) |
Plants larger, leafy stems 10-50 mm long, 2.0-5.0 mm wide; costa percurrent |
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27 Fissidens involutus |
+ |
Plants smaller, leafy stems 3-7 mm long, 2.5-3.0 mm wide; costa usually ending a few cells below leaf apex |
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(51) |
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51 (50) |
Leaf apices narrowly acute or obtuse to obtusely acute |
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(52) |
+ |
Leaf apices widely acute, often mucronate |
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(53) |
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52 (51) |
Apices of dry leaves often enrolled towards the substrate; apical laminal cells 8-14 µm long |
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var. oblongifolius |
+ |
Apices of dry leaves often enrolled away from the substrate; apical laminal cells 5-8 µm long |
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35b var. hyophilus |
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53 (51) |
Leaves lingulate to lanceolate; laminal cells distinctly mammillose, with indistinctly demarcated walls |
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23 Fissidens gymnogynus |
+ |
Leaves ovate-oblong to ovate-lanceolate; laminal cells more or less mammillose, with well demarcated walls |
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37 Fissidens osmundoides |