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28. Otostegia Benth., Lab. Gen. et Sp.  601.  1834.  Benth. in DC., Prodr. 12: 552. 1848; Boiss., Fl. Or. 4: 766. 1879; Hook. f., Fl. Brit. Ind. 4: 679. 1885; Briquet in Engler & Prantl, Nat. Pflanzenfam. ed. 1, 4, 3A: 258. 1896; Kudrjaschev, Monograph of Otostegia, in Trudy Bot. Sekt. Kom. Nauk Uzbek SSSR 1939 [in Russian]; Mukerjee in Rec. Bot. Surv. Ind. 14, 1: 157. 1940; Gorshkova in Komarov, Fl. URSS 21: 182. 1954; Hedge & Lamond in Notes Roy. Bot. Gard. Edinb. 28: 125, f. 3. 1968; Rech. f., Fl. Iran. 150: 347. 1982.  
I.C. Hedge
 
 
 
 
Shrubs or subshrubs, glabrous or pubescent with simple hairs. Leaves not divided, entire, crenate or dentate, sessile or shortly petiolate, spine-tipped or not; hard spines present in leaf axils On ours). Verticillasters distant, borne in upper leaf axils, 2-6(-8)-flowered. Calyx clearly differentiated into a tube and a spreading limb, 10-nerved, bilabiate, irregularly 5-toothed or more; in fruit coriaceous with the limb enlarging and horizontally spreading or reflexed. Corolla clearly bilabiate, yellowish-white; tube included within calyx, piloseannulate within; upper lip ± falcate or straight, vinous; lower lip spreading, 3-lobed median lobe larger than laterals. Stamens 4, didynamous; filaments glabrous or pilose; thecae 2-Jocular, glabrous. Style subequally bifid. Nutlets obovoid, apically obtuse-truncate or truncate. 
A distinct genus of c. 20 species with 2 disjunct areas of concentration: one in the C. Asian-Afghan-Pakistan region; the other in the mountains of east Africa (cf. Hedge & Lamond l.c. fig. 3). Probably allied to Ballota L. 
 
 
 
 
 
 
 
 
                        
                         
		
                          
                          
			
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  | 1 | 
  Leaves obovate to suborbicular, crenate; nervation prominent beneath | 
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  1 Otostegia persica | 
 
  | + | 
  Leaves narrowly elliptic - lanceolate, entire; nervation indistinct | 
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  (2) | 
 
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  | 2 (1) | 
  Leaves acuminate and spinulose at apex; corolla white | 
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  2 Otostegia aucheri | 
 
  | + | 
  Leaves obtuse and not spinulose at apex; corolla yellow | 
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  3 Otostegia limbata | 
 
 
 
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Lower Taxa 
 
 
Related Synonym(s): 
 
  
                        
                          
                        
                        
                      
 
                      
	 
                      
                         
		
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