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BFNA | Family List | BFNA Vol. 2 | Bartramiaceae

Bartramia Hedwig, Sp. Musc. 164. 1801.
[conserved name; for the Pennsylvania colonist, John Bartram]

D. Griffin, III

Plants small to robust, in loose to dense, soft or rigid tufts, dull green to glaucous, sometimes yellowish or yellowish brown above, radiculose proximally. Stems erect, often forked, 1--15 cm, roundish in cross section, cortex of somewhat firm walled cells forming an indistinct to distinct hyaloderm. Leaves erect-appressed to spreading or circinate, occasionally flexuose when dry, spreading when moist, slenderly lanceolate-subulate from a more or less sheathing, non plicate base; lamina 2- or 3-stratose at margins or throughout; margins revolute at shoulders, serrate to serrulate distally, teeth single or paired; costa usually strong and abaxially prominent, sometimes obscure in distal acumen and low in profile, subpercurrent to excurrent; distal laminal cells small, subquadrate to oblong linear, firm-walled, prorulose on both surfaces; basal cells elongate to rectangular or linear, thin-walled or infrequently thick walled toward costa, pale, smooth. Sexual condition dioicous, autoicous or synoicous; perigonia gemmiform; perichaetial leaves little differentiated, typically less prorulose and with a laxer areolation than in stem leaves. Seta elongate, rarely short, erect or sometimes curved, smooth. Capsule usually inclined and asymmetric, less frequently erect and symmetric, subglobose, oblique mouthed, furrowed or rarely wrinkled when dry; operculum convex with a low, blunt beak; annulus not distinct; peristome double, single or lacking; teeth reddish brown, without intermediate thickenings on back, smooth or papillose, sometimes cleft above the middle; endostome yellowish, sometimes lacking, segments keeled, cilia rudimentary or none. Spores reniform to subspherical, papillose, 15--40 µm.

Species 90 (7 in the flora). Worldwide (except Antarctica), and especially diverse in montane tropics.

Bartramia is generally recognized by the linear-lanceolate leaves and sheathing base with a distal lamina that is 2- or 3-stratose at the margins or throughout. Bartramia stricta lacks the sheathing leaf base but otherwise accords with the generic concept.


Crum, H. A. and L. E. Anderson 1981. Bartramiaceae. In: Mosses of Eastern North America, vols 1--2. New York. Flowers, S. 1935. Bartramia. In: A. J. Grout, Moss Fl. N. Amer. Vol. II, part 3, pp. 157--163. Newfane, Vermont. Flowers, S. 1973. Mosses: Utah and the West. Brigham Young University Press. Provo. Fransén, S. 1995. A taxonomic revision of Neotropical Bartramia section Vaginella C. Muell. Lindbergia 20: 147-179. Ireland, R. R. 1982. Moss Flora of the Maritime Provinces. Publ. Bot. No. 13, National Museums of Canada, Ottawa. Lawton, E. 1971. Moss Flora of the Pacific Northwest. Hattori Botanical Laboratory. Nichinan.

1 Leaves large, linear, 4--7 mm, not strict, base not sharply distinct; plants large, to 10 cm.   (2)
+ Leaves smaller, subulate, usually abruptly narrowed and divergent from a hyaline, sheathing base (except B. stricta), usually strict, 2.5--5 mm; plants small, to 5 cm.   (3)
2 (1) Seta short, equaling or slightly longer than capsule.   Bartramia halleriana
+ Seta many times longer than capsule.   Bartramia pomiformis
3 (1) Leaf base not sheathing, not abruptly distinct.   Bartramia stricta
+ Leaf base sheathing, hyaline, abruptly distinct from distal lamina.   (4)
4 (3) Costa in distal limb obscure; distal laminal cells 25--45 µm, prorulae low.   Bartramia ithyphylla
+ Costa in distal limb prominent; distal laminal cells 10--30 µm, prorulae high.   (5)
5 (4) Shoulders delicate, often eroded; leaf tips fragile, usually broken.   Bartramia potosica
+ Shoulders relatively firm, not eroded; leaf tips mostly intact.   (6)
6 (5) Basal cells of leaf differentiated with thick walled cells toward costa and thin-walled cells toward margin; leaf margins revolute distal to shoulders.   Bartramia brevifolia
+ Basal cells of leaf thin-walled throughout; leaf margins typically plane distal to shoulders.   Bartramia subulata

Lower Taxa

Related Synonym(s):


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