227. Tephroseris (Reichenbach) Reichenbach, Deut. Bot. Herb.-Buch. 87. 1841.
[Greek tephros, ashlike or ash-colored, and seris, endive or chicory, presumably alluding to color of the densely woolly leaves]
Theodore M. Barkley†, David F. Murray
Cineraria Linnaeus [unranked] Tephroseris Reichenbach in J. C. Mössler and H. G. L. Reichenbach, Handb. Gewächsk. ed. 2, 2: 1498. 1829
Annuals, biennials, or perennials, (5–)10–100+ cm (rhizomatous or caudices ± erect; plants usually arachnose, floccose, lanate, tomentose, or villous, sometimes unevenly glabrate). Stems 1 or more (loosely clustered), erect. Leaves basal and cauline; alternate; petiolate (basal and proximal cauline; distal leaves usually sessile, smaller, bractlike); blades pinnately nerved, lanceolate, linear-oblanceolate, oblanceolate, ovate, or subrhombic (bases tapering or contracted to petioles), margins entire or dentate, denticulate, subentire, subpinnatifid, or wavy, faces usually arachnose, floccose, lanate, tomentose, or villous, sometimes unevenly glabrate. Heads radiate or discoid, borne singly or (2–40+) in corymbiform arrays. Calyculi 0. Involucres hemispheric or campanulate to turbinate, 8–12+ mm diam. Phyllaries persistent, usually 8, 13, or 21 in (1–)2 series, erect, distinct (margins interlocking), lance-linear to lanceolate or oblong, equal, margins ± scarious (abaxial faces usually arachnose, floccose, lanate, tomentose, or villous, sometimes unevenly glabrate). Receptacles flat or ± dome-shaped (not conic), smooth, epaleate. Ray florets 0 or mostly 8, 13, or 21, pistillate, fertile; corollas usually yellow, orange, or orange-yellow, sometimes ochroleucous or white [brick-colored, purplish] (laminae usually 5–20 mm, sometimes 1–3 mm). Disc florets 30–80+, bisexual, fertile; corollas usually yellow, orange, or orange-yellow, sometimes ochroleucous or white [brick-colored, purplish], tubes longer than or equaling campanulate throats, lobes 5, erect or recurved, lance-linear (anther collars cylindric); style branches: stigmatic areas continuous, apices rounded-truncate. Cypselae ± cylindric, 10-ribbed or -nerved, glabrous or puberulent; pappi persistent, of 30–60+, white, whitish, or brownish, barbellulate bristles (equaling or slightly exceeding involucres, sometimes exceeding involucres to 10 mm in T. palustris). x = 24.
Species 40–50 (6 in the flora): n North America, centered in n Eurasia.
Species of Tephroseris are variable and poorly defined; their nomenclature is complex. The present treatment is provisional. Tephroseris has been treated within Senecio in most North American floristic studies (T. M. Barkley 1999). Moreover, the tradition has been to treat much of the variation as varieties or subspecies within a broadly circumscribed Senecio (Tephroseris) atropurpureus (e.g., Barkley 1978; E. Hultén 1968; H. J. Scoggan 1978–1979, part 4; S. L. Welsh 1974); current thought is to recognize more species and thereby bring North American species concepts more into accord with those of Russian botanists (e.g., S. S. Kharkevich 1992, vol. 6; I. M. Krasnoborov 1997, vol. 13; E. Wiebe 2000). The circumscription of Tephroseris was discussed by B. Nordenstam (1978). The Eurasian Tephroseris atropurpureus (Ledebour) B. Fedtschenko, in the strict sense, and T. subfrigida (Komarov) Holub may occur in far western Alaska. The former resembles T. frigida; it has smaller heads, narrower, purplish phyllaries, and a different "aspect." Tephroseris subfrigida is a relatively tall, thin plant with phyllaries purplish on the distal one-third; the bases of the heads have a light yellowish, non-woolly tomentum.
SELECTED REFERENCE
Cufodontis, G. 1933. Kritische Revision von Senecio Section Tephroseris. Feddes Repert. Spec. Nov. Regni Veg. 70: 1–266.
