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FOC | Family List | FOC Vol. 6 | Ranunculaceae

6. Cimicifuga Wernischeck, Gen. Pl. 298. 1763.

升麻属 sheng ma shu

Li Liangqian; Anthony R. Brach

Herbs perennial. Rhizome robust, creeping, with fibrous roots. Stem terete, apically usually several branched. Leaves 1--3 × ternately sect or subpinnately compound, long petiolate. Inflorescence densely racemose, sometimes spicate, simple or branched; rachis densely glandular pubescent and hairy; bracts subulate to narrowly triangular, small. Flowers dense, small, actinomorphic, hermaphroditic or rarely unisexual and plants dioecious. Sepals 4 or 5, petaloid, white, obovate-orbicular, caducous. Petals elliptic to suborbicular, entire, slightly concave or forked-lobed with 2 empty anthers, rarely with a nectary. Stamens numerous; filaments narrowly linear to filamentous; anthers yellow, broadly ellipsoid to suborbicular. Follicles 1--8, stalked or sessile, oblong-ellipsoid to obovate-ellipsoid, adaxially convex veined, apex beaked. Seeds few, yellowish brown, ellipsoid to narrowly ellipsoid.

About 18 species: N temperate regions; eight species (three endemic) in China.

Elsewhere (Compton et al., Taxon 47: 593–634. 1998), Cimicifuga has been transferred to Actaea . However, for the present account, as in FRPS, one of us (Li) prefers to maintain Cimicifuga as distinct.


1 Leaves 1 × ternate, subbasal; inflorescence spicate; carpels 1, sometimes 2   1 C. japonica
+ Leaves 2 or 3 × ternately pinnate; inflorescence racemose; carpels 2--8.   (2)
       
2 (1) Flower unisexual, plants dioecious   8 C. dahurica
+ Flower hermaphroditic.   (3)
       
3 (2) Inflorescence unbranched or with several short branches at base.   (4)
+ Inflorescence (2--)4--20-branched.   (5)
       
4 (3) Plants 1--1.5 m; terminal leaflet ovate or rhombic, 4.5--8.5 × 2--5.5 cm, adaxially glabrous; pedicels 5--8 mm   2 C. simplex
+ Plants 40--90(--140) cm; terminal leaflet ovate or broadly rhombic, 2--3.7 × 1.5--3 cm, adaxially sparsely pubescent; pedicels 1--4 mm   3 C. yunnanensis
       
5 (3) Carpels densely gray pubescent; follicles densely appressed pubescent   7 C. foetida
+ Carpels or follicles glabrous or subglabrous.   (6)
       
6 (5) Terminal leaflet rhombic, papery, abaxially pubescent at veins   6 C. brachycarpa
+ Terminal leaflet ovate or obovate, slightly leathery, abaxially glabrous.   (7)
       
7 (6) Petals entire at apex; terminal leaflet obovate to obovate-elliptic, 4--9 cm wide   4 C. heracleifolia
+ Petals 2-lobed at apex; terminal leaflet ovate, 5.5--14.5 cm wide   5 C. nanchuanensis

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