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19. Cyclea Arnott ex Wight, Ill. Ind. Bot.  1: 22.  1840.  
轮环藤属 lun huan teng shu 
 
 
 
 
Paracyclea Kudô & Yamamoto. 
Vines. Petiole usually long; leaf blade peltate, palmately veined. Inflorescences axillary, terminal or on old stems, thyrsoid, usually narrow, rarely broad and sparse; bracts small. Male flowers: sepals usually 4 or 5(or 6), usually connate, 4- or 5-lobed, rarely free; petals 4 or 5, usually connate, entire or 4-8-lobed, rarely free, sometimes absent; stamens 4 or 5, connate into a peltate synandrium, anthers dehiscing transversely. Female flowers: sepals and petals 1 or 2, opposite to each other, rarely absent; carpel 1, stigma short, 3- or many lobed. Drupes obovate-globose to rotund, often slightly flat, style scar near base; endocarp bony, horseshoe-shaped, abaxially bearing 2 or 3 rows of tubercles on both surfaces of rib; condyle usually 1 or 2 cavities, often perforate between style scar and base. Seed horseshoe-shaped; embryo terete, embedded in endosperm. 
About 29 species: S and SE Asia; 13 species (five endemic) in China. 
 
 
 
 
 
 
 
 
                        
                         
		
                          
                          
			
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  | 1 | 
  Inflorescences cauliflorous or on leafless old branches, broad and large, many branched. | 
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  13 C. polypetala | 
 
  | + | 
  Inflorescences axillary on normal leafy shoots, narrow and few branched (except C. barbata) | 
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  (2) | 
 
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  | 2 (1) | 
  Male inflorescences broad and large, many branched. | 
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  1 C. barbata | 
 
  | + | 
  Male inflorescences narrow, thyrsoid, few branched, sometimes spikelike | 
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  (3) | 
 
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  | 3 (2) | 
  Male flowers with sepals conspicuously connate | 
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  (4) | 
 
  | + | 
  Male flowers with sepals free or only connate at base | 
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  (7) | 
 
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  | 4 (3) | 
  Male sepals forming cup, lobes ca. as long as tube; leaves and branchlets hispid; petioles not peltately inserted. | 
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  10 C. meeboldii | 
 
  | + | 
  Male sepals not forming cup, lobes much shorter than tube; leaves and branchlets puberulent or subglabrous; petioles ± peltately inserted | 
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  (5) | 
 
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    | 
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  | 5 (4) | 
  Male sepals urceolate, pubescent. | 
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  12 C. tonkinensis | 
 
  | + | 
  Male sepals mitriform, glabrous | 
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  (6) | 
 
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  | 6 (5) | 
  Synandrium not exserted; male sepals 2.5-4 mm. | 
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  5 C. racemosa | 
 
  | + | 
  Synandrium exserted; male sepals ca. 2 mm or shorter. | 
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  11 C. debiliflora | 
 
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  | 7 (3) | 
  Rachis and drupes glabrous | 
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  (8) | 
 
  | + | 
  Rachis and drupes pubescent to hispid | 
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  (10) | 
 
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  | 8 (7) | 
  Endocarp ca. 7 mm. | 
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  2 C. sutchuenensis | 
 
  | + | 
  Endocarp less than 5.5 mm | 
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  (9) | 
 
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  | 9 (8) | 
  Endocarp subtruncate or slightly emarginate; leaves 2.5-7 × 1.5-4 cm, glabrous or sparsely pilose, not papillate. | 
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  3 C. hypoglauca | 
 
  | + | 
  Endocarp tapered at base; leaves 6-12 × 3-5.5 cm, abaxially densely setose or papillate. | 
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  4 C. wattii | 
 
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  | 10 (7) | 
  Leaf blade with margins coarsely 1- or 2-toothed. | 
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  9 C. ochiaiana | 
 
  | + | 
  Leaf blade with margins entire or sparsely serrate | 
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  (11) | 
 
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  | 11 (10) | 
  Female flowers with petals as long as sepals; male flowers with petal 1 or absent. | 
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  8 C. gracillima | 
 
  | + | 
  Female flowers with petals much smaller than sepals (female flowers unknown in C. longgangensis) | 
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  (12) | 
 
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  | 12 (11) | 
  Male sepals mitriform, 2.5-4 mm. | 
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  5 C. racemosa | 
 
  | + | 
  Male sepals not mitriform | 
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  (13) | 
 
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  | 13 (12) | 
  Leaf margin long pubescent; male sepals oblong-elliptic. | 
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  6 C. longgangensis | 
 
  | + | 
  Leaf margin shortly puberulent; male sepals obovate or spatulate. | 
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  7 C. insularis | 
 
 
 
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                          List of lower taxa 
 
                        
                          
                        
	
                        
                      
 
                      
	 
                      
                         
		
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