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20. Valerianaceae

败酱科 bai jiang ke

Authors: Deyuan Hong, Fred R. Barrie & Charles D. Bell

Herbs, perennial or biennial, rarely annuals [or shrubs, absent from China], with a characteristic fetid odor caused by monoterpenoid and sesquiterpenoid ethereal oils; vestiture of white, multicellular hairs; taprooted or rhizomatous; roots adventitious. Leaves opposite, basal and cauline, less frequently all basal, simple to pinnatisect or imparipinnate, entire to variously toothed. Inflorescences cymose, simple or compound dichasia or monochasia, bracteate and bracteolate. Flowers perfect or rarely unisexual. Calyx 5-lobed (Nardostachys), reduced to inconspicuous teeth (Patrinia), or setaceous, plumose, and pappuslike (Valeriana). Corolla connate, regular to irregular, sometimes 2-lipped; tube often gibbous (nectariferous) near base; limb (3-)5-lobed; corollas of female flowers 1/3-1/2 size of those of perfect or male flowers. Stamens 1-4, inserted near base of corolla tube, weakly to strongly exserted in bisexual and male flowers, vestigial and included in female flowers; anthers versatile, 4-sporangiate, opening by longitudinal slits. Ovary inferior; carpels 3, 1 fertile, with a solitary, pendulous, apical ovule, 2 sterile; style solitary; stigma entire or 3-lobed. Fruit a cypsela (but commonly referred to as an achene), sometimes winged; sterile locules often reduced; calyx persistent. Seed 1, embryo large, straight, and without endosperm.

Twelve genera and ca. 300 species: nearly cosmopolitan; three genera and 33 species (18 endemic) in China.

Chiu Lien-ching & Wang Han-jin. 1986. Valerianaceae. In: Lu An-ming & Chen Shu-kun, eds., Fl. Reipubl. Popularis Sin. 73(1): 5-44.


1 Stamens 3, rarely 4; calyx segments up to 20, inrolled at anthesis, unrolled in mature fruit, becoming plumose and pappuslike.   3 Valeriana
+ Stamens 4, rarely reduced to 1-3; calyx 5-dentate, not pappuslike in fruit   (2)
       
2 (1) Bracteoles not enlarged into wings in fruit; caudices nearly always covered with fibrous remains of old leaves; leaves entire.   1 Nardostachys
+ Bracteoles enlarged into wings in fruit; caudices without fibrous remains of old leaves; leaves segmented or serrate.   2 Patrinia

Lower Taxa


 

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