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FOC | Family List | FOC Vol. 19 | Valerianaceae

2. Patrinia Jussieu, Ann. Mus. Natl. Hist. Nat. 10: 311. 1807.

败酱属 bai jiang shu

Fedia Adanson; Fuisa Rafinesque.

Herbs, perennial, less frequently biennial; taprooted or rhizomatous; roots or rhizomes strongly stinking. Stems sometimes lignified at base. Basal leaves rosulate, simple to pinnatifid or pinnatisect, often wilted or caducous at anthesis. Cauline leaves opposite, simple to pinnatifid or pinnatisect, margin serrate or dentate, rarely entire. Inflorescence of corymbiform or paniculiform compound dichasia. Flowers 3-6 mm; calyx limb 5-lobed; lobes sinuous, obtusely dentate, ovate or ovate-deltoid, persistent, rarely enlarged in fruit. Corolla yellow, pale yellow, or white, campanulate or funnelform; tube villous within, gibbous basally, nectary sac with dense glands; limb 5-lobed, lobes slightly unequal. Stamens (1-)4, in unequal pairs, inserted at base of corolla tube, usually exserted; longer filaments villous basally, shorter filaments glabrous; anthers oblong, versatile. Style sometimes curved above; stigma capitate or peltate. Achene ovoid or obovoid-oblong; bracteoles reduced or expanded and winglike, detaching as a unit with fruit, 2- or 3-veined, reticulate veins conspicuous. Seeds compressed ellipsoid.

About 20 species: C and E Asia; 11 species (five endemic) in China.

1 Cauline leaves in whorls of 3.   10 P. trifoliata
+ Cauline leaves opposite, decussate   (2)
2 (1) Bracteoles reduced in fruit; peduncles densely hirsute on upper side only; corolla yellow.   6 P. scabiosifolia
+ Bracteoles enlarged and winged in fruit; peduncles uniformly pubescent or hairs in bilateral rows; corolla yellow, pale yellow, or white   (3)
3 (2) Winged bracteoles 2- or occasionally 3-veined   (4)
+ Winged bracteoles usually 3-veined, rarely 2-veined or multiveined   (7)
4 (3) Peduncles hispidulous, hairs less than 0.4 mm; cauline leaves usually pinnatifid, rarely entire   (5)
+ Peduncles hirsute, hairs ca. 1 mm; cauline leaves usually undivided or sometimes with 1-3 pairs of lateral segments   (6)
5 (4) Leaves rigid, leathery, leaf segments obtuse or acute at apex; corolla 6.5-9 mm; bracteoles in fruit 7-9 × 5-7 mm.   7 P. scabra
+ Leaves thin, papery, leaf segments acute to acuminate at apex; corolla 3-4.5 mm; bracteoles in fruit 5.5-6.2 × 4.5-5.5 mm.   2 P. heterophylla
6 (4) Corolla white, limb 3.5-5 mm wide; stamens 4.   11 P. villosa
+ Corolla yellow or pale yellow, very rarely white, limb 2-4 mm wide; stamens 1-4.   4 P. monandra
7 (3) Peduncles scaberulose   (8)
+ Peduncles hirsute   (9)
8 (7) Basal and cauline leaves pinnatisect or bipinnatisect; segments linear.   3 P. intermedia
+ Basal leaves usually shallowly lobed or entire; cauline leaves pinnatifid or pinnatisect, segments various in shape.   5 P. rupestris
9 (7) Plants usually more than 30 cm tall; calyx not obviously enlarged in fruit; leaves glabrous.   1 P. glabrifolia
+ Plants dwarf, less than 25 cm tall; calyx obviously enlarged in fruit; leaves usually pubescent   (10)
10 (9) Rhizomes robust; basal leaves oblanceolate, entire or pinnatifid.   8 P. sibirica
+ Rhizomes usually slender; basal leaves oblanceolate-oblong, all pinnatifid.   9 P. speciosa

  • List of lower taxa


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