1 |
Spikelets 2 per node (at least at middle nodes), rarely 3 or 4 per node |
|
(2) |
+ |
Spikelets 1 per node |
|
(15) |
|
|
|
|
2 (1) |
Glumes much shorter than lemma |
|
(3) |
+ |
Glumes slightly shorter than or equaling lemma |
|
(8) |
|
|
|
|
3 (2) |
Lemma awn more than 10 mm |
|
(4) |
+ |
Lemma awn less than 7 mm |
|
(6) |
|
|
|
|
4 (3) |
Leaf blade flat, 10–15 × 0.5–1 cm. |
|
7 E. sibiricus |
+ |
Leaf blade flat or ± rolled, 3–10 × 0.2–0.5 cm |
|
(5) |
|
|
|
|
5 (4) |
Leaf blade flat, 3–5 mm wide; glume apex with awn 1–4 mm. |
|
8 E. nutans |
+ |
Leaf blade ± rolled, ca. 2 mm wide; glume apex mucronate or acuminate. |
|
9 E. atratus |
|
|
|
|
6 (3) |
Leaf blade 1.5–3 mm wide; glumes oblong, 2–3 mm, 3-veined. |
|
12 E. sinosubmuticus |
+ |
Leaf blade 3–5 mm wide; glumes ovate-lanceolate, oblong-lanceolate, or narrowly lanceolate, 3–4 mm, 1–3-veined |
|
(7) |
|
|
|
|
7 (6) |
Spikelets 13–15 mm, with 4–6 florets; glumes oblong-lanceolate or ovate-lanceolate. |
|
10 E. breviaristatus |
+ |
Spikelets 10–12 mm, with 3 or 4 florets; glumes narrowly lanceolate. |
|
11 E. alpinus |
|
|
|
|
8 (2) |
Rachis densely hirsute, nodes swollen. |
|
6 E. villifer |
+ |
Rachis not densely hirsute, nodes not swollen |
|
(9) |
|
|
|
|
9 (8) |
Culms, leaf blade, and spike all farinaceous; glumes and lemma ± purplish punctate; awn purple, pubescent. |
|
5 E. purpuraristatus |
+ |
Culms, leaf blade, and spike not farinaceous; glumes and lemma not purplish punctate; awn green, glabrous but scabrous |
|
(10) |
|
|
|
|
10 (9) |
Glumes hirsutulous along veins |
|
(11) |
+ |
Glumes scabrous along veins |
|
(12) |
|
|
|
|
11 (10) |
Leaf sheath glabrous; spike purplish; glume apex acuminate or with point ca. 1.5 mm. |
|
2 E. barystachyus |
+ |
Leaf sheath villous at base; spike green; glume apex usually with awn to 5 mm. |
|
4 E. dahuricus |
|
|
|
|
12 (10) |
Glume apex acuminate or with awn 1–4 mm; lemma with usually erect awn (3–)5–13 mm |
|
(13) |
+ |
Glume apex with awn 5–7 mm; lemma with usually reflexed awn 8–20(–40) mm |
|
(14) |
|
|
|
|
13 (12) |
Culms to 120 cm tall, 4- or 5-noded; leaf blade 10–20 × 0.6–1.4 cm; lemma glabrous, or hirsutulous in distal half. |
|
1 E. tangutorum |
+ |
Culms 40–80 cm tall, 2- or 3-noded; leaf blade 5–12 × ca. 0.5 cm; lemma puberulent throughout |
|
4 E. dahuricus |
|
|
|
|
14 (12) |
Leaf blade 1–1.6 cm wide; spikelets 2 or 3(or 4) per node. |
|
3 E. excelsus |
+ |
Leaf blade 0.5–0.9(–1.2) cm wide; spikelets 2 per node, or 1 per node in proximal and distal parts of spike. |
|
4 E. dahuricus |
|
|
|
|
15 (1) |
Lemma with long, strongly recurved awn at seed maturity; if lemma awn short or nearly absent, or only slightly curved, then palea shorter than lemma, ciliate along keels, and apex rounded or emarginate |
|
(16) |
+ |
Lemma with straight awn or awnless; palea slightly shorter to slightly longer than lemma |
|
(52) |
|
|
|
|
16 (15) |
Glumes usually slightly shorter than or equaling first lemma, 5–7-veined |
|
(17) |
+ |
Glumes much shorter than first lemma, (1–)3–5-veined; if glumes only slightly shorter than first lemma then margin thin, scarious, membranous, or very narrowly hyaline |
|
(25) |
|
|
|
|
17 (16) |
Palea oblong-obovate, much shorter than lemma. |
|
14 E. ciliaris |
+ |
Palea oblong, slightly shorter than or equaling lemma |
|
(18) |
|
|
|
|
18 (17) |
Lemma glabrous throughout. |
|
20 E. tibeticus |
+ |
Lemma scabrous, pubescent, or hispidulous throughout |
|
(19) |
|
|
|
|
19 (18) |
Glumes with thin margin. |
|
15 E. canaliculatus |
+ |
Glumes not with thin margin |
|
(20) |
|
|
|
|
20 (19) |
Lemma abaxially hispidulous or laxly spinulose |
|
(21) |
+ |
Lemma abaxially smooth and glabrous (at middle), scabrous, puberulent, or pubescent, sometimes spinulose along veins |
|
(22) |
|
|
|
|
21 (20) |
Lemma abaxially laxly spinulose, awn 2–2.5 cm; culms 30–50 tall. |
|
16 E. sinoflexuosus |
+ |
Lemma abaxially hispidulous, awn 2.5–4.5 cm, culms 60–100 cm tall. |
|
17 E. gmelinii |
|
|
|
|
22 (20) |
Lemma abaxially scabrous, puberulent, or pubescent |
|
(23) |
+ |
Lemma abaxially smooth and glabrous at middle, otherwise spinulose along veins or ± puberulent |
|
(24) |
|
|
|
|
23 (22) |
Lemma abaxially puberulent; culms 25–30 cm tall. |
|
18 E. zhui |
+ |
Lemma abaxially scabrous or pubescent, margin ciliolate; culms 30–90 cm tall. |
|
19 E. formosanus |
|
|
|
|
24 (22) |
Leaf blade usually flat, thin; glume apex acuminate. |
|
21 E. nakaii |
+ |
Leaf blade involute, stiff; glume apex acute or with point 1–5 mm. |
|
22 E. strictus |
|
|
|
|
25 (16) |
Glumes slightly shorter than first lemma, margin thin, scarious, membranous, or very narrowly hyaline |
|
(26) |
+ |
Glumes much shorter than first lemma, margin herbaceous |
|
(35) |
|
|
|
|
26 (25) |
Glume margin scarious |
|
(27) |
+ |
Glume margin thin, membranous, or very narrowly hyaline |
|
(28) |
|
|
|
|
27 (26) |
Culms 80–150 cm tall. |
|
23 E. caninus |
+ |
Culms 30–60 cm tall. |
|
24 E. caesifolius |
|
|
|
|
28 (26) |
Glume margin thin; spikelets pedicellate, pedicel 0.8–9 mm |
|
(29) |
+ |
Glume margin membranous or very narrowly hyaline; spikelets sessile or subsessile |
|
(30) |
|
|
|
|
29 (28) |
Spikelet pedicel 0.8–1.5(–2) mm. |
|
13 E. cacuminis |
|
|
|
|
30 (28) |
Glume margin very narrowly hyaline |
|
(31) |
+ |
Glume margin membranous |
|
(32) |
|
|
|
|
31 (30) |
Culms 50–100 cm tall, 3–5-noded; leaf blade flat, 2.5–8 mm wide. |
|
26 E. antiquus |
+ |
Culms 10–60 cm tall, 2- or 3-noded; leaf blade tightly involute, 1–4 mm wide. |
|
27 E. burchan-buddae |
|
|
|
|
32 (30) |
Lemma glabrous throughout |
|
(33) |
+ |
Lemma puberulent or ± hispidulous |
|
(34) |
|
|
|
|
33 (32) |
Glume apex with short awn; leaf blade adaxial surface villous. |
|
28 E. serpentinus |
+ |
Glume apex awnless, usually acute; leaf blade adaxial surface usually scabrous. |
|
29 E. glaberrimus |
|
|
|
|
34 (32) |
Spikelets with 3 or 4 florets; lemma puberulent. |
|
30 E. pseudocaninus |
+ |
Spikelets with 4 or 5(or 6) florets; lemma hispidulous distally and near margin. |
|
31 E. sinkiangensis |
|
|
|
|
35 (25) |
Glume apex usually tapering into awn 2–7 mm; palea equaling lemma |
|
(36) |
+ |
Glume apex acute, mucronate, or with awn less than 2 mm; palea slightly shorter to slightly longer than lemma |
|
(43) |
|
|
|
|
36 (35) |
Lemma glabrous throughout |
|
(37) |
+ |
Lemma hirsute or hirsutulous throughout, pubescent only at base and margin, or puberulent proximally and hispidulous distally |
|
(39) |
|
|
|
|
37 (36) |
Culms 15–20 cm tall; first lemma 7–9 mm; anthers yellow or purple, 1.2–2.5 mm. |
|
32 E. jacquemontii |
+ |
Culms (15–)40–50 cm tall; first lemma 10–12 mm; anthers black or brown, 2–5 mm |
|
(38) |
|
|
|
|
38 (37) |
First lemma ca. 10 mm; anthers black, ca. 2 mm. |
|
33 E. aristiglumis |
+ |
First lemma 11–12 mm; anthers brown, 4–5 mm. |
|
34 E. shouliangiae |
|
|
|
|
39 (36) |
Lemma pubescent only at base and margin. |
|
37 E. altissimus |
+ |
Lemma hirsute or hirsutulous throughout, or puberulent proximally and hispidulous distally |
|
(40) |
|
|
|
|
40 (39) |
Lemma hirsutulous throughout |
|
(41) |
+ |
Lemma hirsute throughout, or puberulent proximally and hispidulous distally |
|
(42) |
|
|
|
|
41 (40) |
Culms 1- or 2-noded; spikelets with 2 or 3 florets. |
|
33 E. aristiglumis |
+ |
Culms 3-noded; spikelets with 4 or 5 florets. |
|
35 E. confusus |
|
|
|
|
42 (40) |
Lemma hirsute throughout; culms 60–75 cm tall; palea ciliolate along keels distally. |
|
36 E. anthosachnoides |
+ |
Lemma puberulent proximally, hispidulous distally; culms 20–45 cm tall; palea puberulent between keels distally. |
|
39 E. serotinus |
|
|
|
|
43 (35) |
Palea equaling or slightly longer than lemma; anthers 2.5–3 mm; lemma awn (2.5–)3.5–5.5 cm |
|
(44) |
+ |
Palea slightly shorter than lemma; anthers 1–2 mm; lemma awn 1–3 cm |
|
(45) |
|
|
|
|
44 (43) |
Leaf blade 1–2 mm wide; palea ciliate along keels. |
|
40 E. retroflexus |
+ |
Leaf blade 2–4 mm wide; palea glabrous along keels. |
|
41 E. leiotropis |
|
|
|
|
45 (43) |
Spike somewhat nitid; lemma glabrous or sparsely puberulent abaxially. |
|
42 E. brevipes |
+ |
Spike not nitid; lemma pubescent, hirsutulous, or scabrous abaxially |
|
(46) |
|
|
|
|
46 (45) |
Spikelets sessile or very shortly pedicellate, with 1–4(or 5) florets |
|
(47) |
+ |
Spikelets usually sessile, with 4–9 florets |
|
(49) |
|
|
|
|
47 (46) |
Spikelets usually very shortly pedicellate, with 3 or 4(or 5) florets. |
|
43 E. schrenkianus |
+ |
Spikelets sessile; with 1–3(–5) florets |
|
(48) |
|
|
|
|
48 (47) |
Spikelets with 1–3 florets; culms 70–150 cm tall, 5–7-noded. |
|
37 E. altissimus |
+ |
Spikelets with 2–5 florets; culms 40–80 cm tall, 3- or 4-noded. |
|
38 E. laxinodis |
|
|
|
|
49 (46) |
Lemma hirsute throughout, especially distally and near margin, first lemma 13–14 mm. |
|
36 E. anthosachnoides |
+ |
Lemma ± hirsutulous or scabrous abaxially, first lemma 9–11 mm |
|
(50) |
|
|
|
|
50 (49) |
Lemma ± hirsutulous abaxially, first lemma ca. 11 mm, awn 3–4 cm; spikelets with 7–9 florets. |
|
44 E. pulanensis |
+ |
Lemma scabrous abaxially, rarely hirsutulous, first lemma 9–11 mm, awn 1.8–3 cm; spikelets with 4–7 florets |
|
(51) |
|
|
|
|
51 (50) |
Spike ± purplish, nodding, 13–15 cm; glumes oblong-lanceolate, apex acute; lemma scabrous or hirsutulous abaxially. |
|
45 E. purpurascens |
+ |
Spike usually glaucous green, slender and flexuose, 7–9 cm; glumes linear-lanceolate, apex usually acuminate; lemma scabrous abaxially. |
|
46 E. tschimganicus |
|
|
|
|
52 (15) |
Lemma awn shorter than 7 mm or absent |
|
(53) |
+ |
Lemma awn more than 7 mm (but 1–35 mm E. abolinii) |
|
(68) |
|
|
|
|
53 (52) |
Lemma apex truncate, with 3 teeth. |
|
51 E. tridentatus |
+ |
Lemma ± narrowed toward apex, without teeth |
|
(54) |
|
|
|
|
54 (53) |
Palea equaling or slightly longer than lemma, apex narrowly truncate |
|
(55) |
+ |
Palea slightly shorter to longer than lemma, apex emarginate, rounded, or subrounded |
|
(58) |
|
|
|
|
55 (54) |
Lemma ± pilose. |
|
47 E. mutabilis |
+ |
Lemma smooth, glabrous |
|
(56) |
|
|
|
|
56 (55) |
Plants without rhizomes. |
|
48 E. magnicaespes |
+ |
Plants ± with rhizomes |
|
(57) |
|
|
|
|
57 (56) |
First lemma ca. 10 mm, callus smooth, glabrous; anthers ca. 3 mm. |
|
49 E. alashanicus |
+ |
First lemma 7.8–9 mm, callus puberulent; anthers 4.5–5 mm. |
|
52 E. elytrigioides |
|
|
|
|
58 (54) |
Glumes densely pubescent adaxially. |
|
50 E. jufinshanicus |
+ |
Glumes not as above |
|
(59) |
|
|
|
|
59 (58) |
Palea 2/3–3/4 as long as lemma, subsmooth along keels. |
|
53 E. grandis |
+ |
Palea equaling, subequaling, or longer than lemma, ± ciliate along keels |
|
(60) |
|
|
|
|
60 (59) |
Glumes 4–9-veined; palea slightly shorter to slightly longer than lemma |
|
(61) |
+ |
Glumes 3–5-veined; palea equaling lemma |
|
(64) |
|
|
|
|
61 (60) |
Culms 30–80 cm tall; leaf blade 2–4 mm wide |
|
(62) |
+ |
Culms usually more than 100 cm tall; leaf blade 5–11 mm wide |
|
(63) |
|
|
|
|
62 (61) |
Spikelets 1.4–1.8 cm, with 5 or 6 florets; lemma margin broadly membranous. |
|
59 E. viridulus |
+ |
Spikelets 1–1.4 cm, with 2–4 florets; lemma margin not membranous. |
|
60 E. cheniae |
|
|
|
|
63 (61) |
Spikelets with 3–5 florets; glumes 5–7-veined; culms 100–160 cm tall. |
|
61 E. intramongolicus |
+ |
Spikelets with 5–9 florets; glumes 7–9-veined; culms 50–120 cm tall. |
|
62 E. nevskii |
|
|
|
|
64 (60) |
Spikelets with 2 or 3 florets. |
|
55 E. kronokensis |
+ |
Spikelets with 3–6 florets |
|
(65) |
|
|
|
|
65 (64) |
Culms ca. 100 cm tall, 4- or 5-noded. |
|
54 E. sylvaticus |
+ |
Culms 20–60 cm tall, 2–4-noded |
|
(66) |
|
|
|
|
66 (65) |
Culms 45–60 cm tall, 2- or 3-noded. |
|
56 E. yushuensis |
+ |
Culms 20–45 cm tall, 2–4-noded |
|
(67) |
|
|
|
|
67 (66) |
Culms ca. 45 cm tall, usually 4-noded, palea keels wingless and ciliolate. |
|
57 E. leianthus |
+ |
Culms 20–25 cm tall, usually 2-noded; palea keels winged and ciliate. |
|
58 E. humilis |
|
|
|
|
68 (52) |
Glumes 5–7(–9)-veined, slightly shorter or longer than lemma |
|
(69) |
+ |
Glumes (2 or)3–5-veined, at least proximal glume 3-veined, usually much shorter than lemma, or if subequaling lemma then apex shortly awned |
|
(76) |
|
|
|
|
69 (68) |
Glumes (especially distal one) longer than lemma |
|
(70) |
+ |
Glumes (especially proximal one) slightly shorter than lemma |
|
(72) |
|
|
|
|
70 (69) |
Spike somewhat lax; palea shorter than lemma; anthers 3–7 mm. |
|
81 E. abolinii |
+ |
Spike dense, secund; palea subequaling lemma; anthers 2.5–3 mm |
|
(71) |
|
|
|
|
71 (70) |
Glume margin membranous; lemma scabrous or appressed setose. |
|
82 E. fedtschenkoi |
+ |
Glume margin not membranous; lemma pubescent. |
|
83 E. platyphyllus |
|
|
|
|
72 (69) |
Spikelets pedicellate, pedicel 1–2.5 mm. |
|
85 E. magnipodus |
+ |
Spikelets sessile |
|
(73) |
|
|
|
|
73 (72) |
Lemma ± glabrous abaxially, margin narrowly membranous. |
|
84 E. shandongensis |
+ |
Lemma puberulent, pilose, or ± hirsutulous abaxially, margin not membranous |
|
(74) |
|
|
|
|
74 (73) |
Lemma margin ciliate. |
|
86 E. pendulinus |
+ |
Lemma margin not ciliate |
|
(75) |
|
|
|
|
75 (74) |
Glume margin scarious; leaf blade abaxial surface glabrous or scabrous, adaxial surface scabrous or pubescent. |
|
87 E. komarovii |
+ |
Glume margin not scarious; leaf blade both surfaces pilose or scabrous. |
|
88 E. tianschanigenus |
|
|
|
|
76 (68) |
Glumes always and lemma sometimes with membranous or scarious margin |
|
(77) |
+ |
Glumes and lemma without membranous or scarious margin |
|
(85) |
|
|
|
|
77 (76) |
Glumes and lemma with membranous or scarious margin |
|
(78) |
+ |
Glumes with membranous margin; lemma without such margin |
|
(79) |
|
|
|
|
78 (77) |
Lemma glabrous to hispidulous, margin membranous. |
|
63 E. kamoji |
+ |
Lemma hirsutulous proximally and distally, subglabrous at middle, long ciliate near margin, margin broadly scarious. |
|
64 E. hybridus |
|
|
|
|
79 (77) |
Culms 50–60 cm tall, 2- or 3-noded |
|
(80) |
+ |
Culms 60–100 cm tall, 4–7-noded |
|
(81) |
|
|
|
|
80 (79) |
Glumes 5–6 mm; lemma awn 5–10 mm; anthers black. |
|
65 E. curtiaristatus |
+ |
Glumes 9–12 mm; lemma awn 12–15 mm; anthers yellow. |
|
66 E. scabridulus |
|
|
|
|
81 (79) |
Spikelets 3.5–4.5 cm, with 5–10 florets. |
|
67 E. yangiae |
+ |
Spikelets to 2 cm, with 3–9 florets |
|
(82) |
|
|
|
|
82 (81) |
Spikelets ± pedicellate; glumes oblong-lanceolate; palea scabrous along keels distally. |
|
68 E. dolichatherus |
+ |
Spikelets sessile; glumes narrowly lanceolate; palea ± ciliate along keels |
|
(83) |
|
|
|
|
83 (82) |
Glumes and lemma usually smooth, glabrous, rarely scabrous along veins. |
|
70 E. calcicola |
+ |
Glumes and lemma hirsute or villous |
|
(84) |
|
|
|
|
84 (83) |
Glumes lanceolate, subequal, 5–6 mm, 2-veined, hispidulous along veins; lemma densely villous abaxially. |
|
71 E. trichospiculus |
+ |
Glumes oblong-lanceolate, hirsute, proximal one 5–6.5 mm, 3- or 4-veined, distal one 6.5–7.5 mm, 4- or 5-veined; lemma usually densely pubescent proximally, scabrous-ciliate distally. |
|
69 E. tenuispicus |
|
|
|
|
85 (76) |
Lemma ± puberulent abaxially |
|
(86) |
+ |
Lemma smooth or scabrous abaxially |
|
(89) |
|
|
|
|
86 (85) |
Spikelets 12–16 mm, with 3–5 florets |
|
(87) |
+ |
Spikelets 9–12 mm, with 2 or 3 florets |
|
(88) |
|
|
|
|
87 (86) |
Glumes oblong-lanceolate, 7–10 mm; lemma awn 10–18 mm. |
|
72 E. sinicus |
+ |
Glumes lanceolate, 2–5 mm; lemma awn 6–8 mm. |
|
74 E. caianus |
|
|
|
|
88 (86) |
Culms usually 2-noded; glume apex acuminate. |
|
76 E. hongyuanensis |
+ |
Culms 5–7-noded; glume apex with awn 1.5–3 mm. |
|
75 E. debilis |
|
|
|
|
89 (85) |
Lemma callus glabrous or subglabrous. |
|
78 E. angustispiculatus |
+ |
Lemma callus distinctly hirsute |
|
(90) |
|
|
|
|
90 (89) |
Spikelets with 2 or 3 florets; lemma callus hairs 1–3 mm. |
|
73 E. puberulus |
+ |
Spikelets with 4–8 florets; lemma callus hairs 4–10 mm |
|
(91) |
|
|
|
|
91 (90) |
Lemma callus hairs 6–10 mm. |
|
77 E. barbicallus |
+ |
Lemma callus hairs 4–6 mm |
|
(92) |
|
|
|
|
92 (91) |
Leaf blade adaxial surface scabrous; lemma scabrous abaxially. |
|
79 E. hondae |
+ |
Leaf blade adaxial surface pubescent; lemma smooth and glabrous abaxially. |
|
80 E. alienus |