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FOC | Family List | FOC Vol. 2-3 | Pteridaceae

19. Haplopteris C. Presl, Tent. Pterid. 141.1836.

书带蕨属 shu dai jue shu

Authors: Zhang Xianchun & Michael G. Gilbert

Epiphytic or epilithic grasslike plants. Fronds simple, linear, entire, glabrous. Venation consisting of a costa and oblique unbranched veins, free except for a submarginal fertile connecting vein. Sori in one elongate marginal or submarginal groove on each side of lamina, immersed, rarely superficial; paraphyses long with a dark obconic head. Spores monolete, most ellipsoid, sometimes fusiform, smooth, transparent.

About 40 unclear species: tropical and subtropical Asia; 13 species in China.

Most of the Old World species formally classified in Vittaria now belong to Haplopteris.

Vittaria suberecta Hayata (Icon. Pl. Formosan. 6: 161. 1916), described from Taiwan, could not be treated here because no material was seen by the present authors.


1 Soral line marginal, immersed in groove, open outward   (2)
+ Soral line submarginal   (3)
       
2 (1) Lamina up to 100 cm or longer, ca. 10 mm wide; stipe long, slender; scales dark brown.   12 H. elongata
+ Lamina 8-30 cm, 2-4 mm wide; stipe short; scales yellow-brown.   13 H. anguste-elongata
       
3 (1) Soral line superficial; lamina margins straight or revolute   (4)
+ Soral line immersed in groove between costa and margins of lamina; lamina margins revolute   (8)
       
4 (3) Scales dark brown, small, straight, areole wall thick; soral line submarginal, covered by revolute margins of lamina.   1 H. amboinensis
+ Scales yellow-brown, large, soft, twisted, areole wall thin; soral line with a broad area between margins of lamina   (5)
       
5 (4) Lamina 2-3 mm wide; scales 3-7 ?ca. 0.2 mm; soral line close to costa.   5 H. mediosora
+ Lamina over 5 mm wide; scales 10-20 ?0.5-2.5 mm; soral line with a broad area between costa   (6)
       
6 (5) Stipe short, thick; lamina 10-30 mm wide; scales 10-20 mm.   2 H. doniana
+ Stipe absent, or slender; lamina 5-12 mm wide; scales ca. 10 mm   (7)
       
7 (6) Lamina 6-12 mm wide; soral line 1-1.5 mm away from margins of lamina.   3 H. taeniophylla
+ Lamina 5-6 mm wide; soral line ca. 1 mm away from margins of lamina.   4 H. himalayensis
       
8 (3) Lamina 5-10 mm wide, or more   (9)
+ Lamina (0.5-)2-4(-11) mm wide   (10)
       
9 (8) Costa not raised adaxially, lamina with a narrow concave line adaxially.   6 H. plurisulcata
+ Costa raised adaxially, lamina with a broad concave line on each side of costa adaxially.   7 H. fudzinoi
       
10 (8) Scales dark or light brown, 5-10 mm   (11)
+ Scales yellow-brown, 2-3 mm   (12)
       
11 (10) Scales light brown; lamina with soral line very close to costa.   8 H. linearifolia
+ Scales dark brown; lamina with soral line close to margins.   9 H. hainanensis
       
12 (10) Costa flattened abaxially, wide; soral line covered by costa.   10 H. sikkimensis
+ Costa raised abaxially, narrow; soral line with a broad area between costa.   11 H. flexuosa

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