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FOC | Family List | FOC Vol. 15 | Primulaceae

1. Lysimachia Linnaeus, Sp. Pl. 1: 146. 1753.

珍珠菜属 zhen zhu cai shu

Herbs erect or procumbent, rarely suffruticose, glabrous or pubescent, often with internal glands appearing as pustulate dots or stripes. Leaves alternate, opposite, or whorled, usually entire. Flowers solitary in axils of upper leaves or in terminal and axillary panicles or racemes, often shortened into capitate clusters, with bracts. Calyx usually 5(or rarely 6--9)-parted. Corolla white or yellow, rarely pink, homomorphic, rarely heteromorphic, subrotate or campanulate, deeply 5(or rarely 6--9)-parted; lobes contorted in bud. Filaments free or connate into a ring or tube at base and ± adnate to corolla tube; anthers basifixed, dorsifixed or versatile opening by apical pores or by lateral slits. Capsule subglobose, usually dehiscing by valves, rarely indehiscent.

Approximately 180 species in temperate and subtropical parts of the northern hemisphere, but with a few species in Africa, Australia, and South America; 138 species occur in China.


Key 1

1 Stems 1-1.5 cm tall; flowers forming terminal racemes on leafless scapes   1 Lysimachia scapiflora
+ Stems more than 20 cm tall; flowers solitary or in racemes on leafy stems.   (2)
       
2 (1) Leaves terminal.   (3)
+ Leaves alternate or opposite, not terminal.   (7)
       
3 (2) Flowers in many-flowered racemes, 6-9 cm; capsules indehiscent   2 Lysimachia insignis
+ Flowers in clusters or extremely shortened racemes; capsules dehiscing by valves.   (4)
       
4 (3) Leaf blade rounded to obtuse at apex; calyx lanceolate, 6-8 mm; corolla lobes acute at apex   6 Lysimachia crassifolia
+ Leaf blade acute at apex; calyx ovate, 2-4 mm; corolla lobes obtuse at apex.   (5)
       
5 (4) Inflorescences distributed along stems under leaf whorls   3 Lysimachia saxicola
+ Inflorescences subterminal.   (6)
       
6 (5) Leaves nearly equal, rounded at base   4 Lysimachia dushanensis
+ Leaves unequal, cordate at base   5 Lysimachia linguiensis
       
7 (2) Leaves opposite.   (8)
+ Leaves alternate.   (10)
       
8 (7) Leaf blade cuneate to rounded at base   36 Lysimachia engleri
+ Leaf blade cordate at base.   (9)
       
9 (8) Flowers solitary; leaf blade 1-4 cm   37 Lysimachia cordifolia
+ Flowers usually in pairs; leaf blade 4.5-8 cm   38 Lysimachia biflora
       
10 (7) Anthers conical, less than 3.5 × as long as wide; stems herbaceous.   (11)
+ Anthers linear, more than 4 × as long as wide; stems subwoody.   (21)
       
11 (10) Flowers in racemes.   (12)
+ Flowers solitary, axillary.   (15)
       
12 (11) Bracts subulate; leaf blade with more than 10 pairs of lateral veins   25 Lysimachia physaloides
+ Bracts leaflike; leaf blade with 5-7 pairs of lateral veins.   (13)
       
13 (12) Stems not winged; flowers subverticillate   28 Lysimachia subverticillata
+ Stems winged; flowers racemose.   (14)
       
14 (13) Racemes corymbiform, 2-5-flowered   26 Lysimachia wilsonii
+ Racemes elongated, 5- to many flowered   27 Lysimachia nutantiflora
       
15 (11) Stems ± procumbent at base, upper part erect, rarely branched; leaf blade broadly ovate to elliptic.   (16)
+ Stems erect or sometimes arcuate at base, usually branched; leaf blade lanceolate to narrowly elliptic or oblong-lanceolate.   (18)
       
16 (15) Calyx 7-12 mm; corolla 2-3.5 cm in diam.; leaf blade attenuate to cuneate at base   29 Lysimachia foenum-graecum
+ Calyx 2.5-6 mm; corolla 1.2-2 cm in diam.; leaf blade rounded or slightly cordate, rarely cuneate at base.   (17)
       
17 (16) Corolla 1.5-2 cm in diam.; calyx 4-6 mm; plants not aromatic   30 Lysimachia trichopoda
+ Corolla ca. 1.2 cm in diam; calyx ca. 2.5 mm; plants aromatic when dry   31 Lysimachia pseudotrichopoda
       
18 (15) Corolla 3-5 mm; calyx lobes narrowly ovate, ca. as long as or slightly shorter than corolla; capsules 3.5-4 mm in diam.   32 Lysimachia brachyandra
+ Corolla more than 6 mm; calyx lobes ovate to ovate-elliptic, 1/3 to 1/2 × as long as corolla; capsules more than 5 mm in diam.   (19)
       
19 (18) Flowers in racemes continuing apically into vegetative shoots   35 Lysimachia cauliflora
+ Flowers solitary, in axils of upper leaves.   (20)
       
20 (19) Corolla 10-15 mm; anthers opening by apical pores   33 Lysimachia grandiflora
+ Corolla 6-8 mm; anthers opening by lateral slits   34 Lysimachia laxa
       
21 (10) Flowers in elongated racemes.   (22)
+ Flowers solitary or 2 to many flowers crowded in leaf axils.   (24)
       
22 (21) Petiole less than 2 mm; leaf blade rounded or subcordate at base; bracts subulate   7 Lysimachia filipes
+ Petiole 5-30 mm; leaf blade subattenuate at base; at least lower bracts leaflike.   (23)
       
23 (22) Calyx lobes lanceolate   8 Lysimachia heterobotrys
+ Calyx lobes ovate   9 Lysimachia chapaensis
       
24 (21) Pedicel 3-10(-12) mm.   (25)
+ Pedicel usually more than 12 mm.   (27)
       
25 (24) Leaf blade 6-12 × 3-7 cm; calyx 6-9 mm   10 Lysimachia inaperta
+ Leaf blade 0.6-1.7 × 0.2-0.8 cm; calyx 3-4 mm.   (26)
       
26 (25) Leaves sparsely setulose adaxially; corolla epunctate   11 Lysimachia aspera
+ Leaves glabrous; corolla glandular punctate   12 Lysimachia tianyangensis
       
27 (24) Leaf blade linear or leathery, veinlets obscure.   (28)
+ Leaf blade neither linear nor leathery, veinlets conspicuous.   (30)
       
28 (27) Leaf blade linear, subfalcate at apex, herbaceous   13 Lysimachia vittiformis
+ Leaf blade neither linear nor falcate at apex, subleathery.   (29)
       
29 (28) Leaves oblanceolate, ± crowded in interrupted whorls   14 Lysimachia pittosporoides
+ Leaves lanceolate, evenly distributed along stems   15 Lysimachia millietii
       
30 (27) Calyx ca. as long as to slightly longer than corolla   16 Lysimachia evalvis
+ Calyx shorter than corolla.   (31)
       
31 (30) Stems conspicuously angular, usually herbaceous.   (32)
+ Stems terete, subwoody.   (34)
       
32 (31) Calyx 5-6 mm, lobes subulate; leaf blade truncate to subcordate at base   22 Lysimachia jingdongensis
+ Calyx 2-4 mm, lobes acuminate; leaf blade subattenuate at base.   (33)
       
33 (32) Stems angular or winged, usually much branched; leaf blade ovate to lanceolate   23 Lysimachia capillipes
+ Stems angular, usually simple; leaf blade lanceolate   24 Lysimachia lancifolia
       
34 (31) Pedicel ca. as long as to shorter than petiole; capsules indehiscent   17 Lysimachia navillei
+ Pedicel much longer than petiole; capsules dehiscing by valves.   (35)
       
35 (34) Capsule 3-4 mm.   (36)
+ Capsule 5-6 mm.   (37)
       
36 (35) Leaf blade elliptic-lanceolate to lanceolate; free parts of filaments distinctly longer than sinus of cordate base of anthers   18 Lysimachia fooningensis
+ Leaf blade ovate to ovate-lanceolate; free parts of filaments ca. as long as sinus of cordate base of anthers   19 Lysimachia microcarpa
       
37 (35) Leaf blade elliptic to elliptic-lanceolate, 1-3(-4) cm wide   20 Lysimachia ardisioides
+ Leaf blade ovate to elliptic or broadly elliptic, 1.3-7(-9) cm wide   21 Lysimachia petelotii

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