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FOC | Family List | FOC Vol. 23 | Cyperaceae

10. Schoenoplectus (Reichenbach) Palla, Verh. K. K. Zool.-Bot. Ges. Wien. 38 (Sitzungsber.): 49. 1888.

水葱属 shui cong shu

Authors: Prof. Song-Yun Liang, Gordon C. Tucker & Tetsuo Koyama

Scirpus subg. Schoenoplectus Reichenbach, Icon. Fl. Germ. Helv. 8: 40. 1846; Schoenoplectiella Lye.

Herbs, annual or perennial, sometimes with creeping rhizomes. Culms tufted, not nodose above base. Leaves usually reduced to a sheath or more rarely developing a ligulate blade. Involucral bracts culmlike, erect. Inflorescence a pseudolateral anthela or more frequently capitate with (1 or)few to many spikelets. Spikelets ovoid to ellipsoid. Glumes many, spiral (rarely pseudodistichous), deciduous or rarely persistent, often each subtending a bisexual flower. Perianth bristles 0-6, needlelike, shorter to longer than nutlet, smooth, scabrous, or plumose, deciduous with nutlet. Stamens 1-3. Style base not distinct, not thickened, persistent. Nutlet obovoid, ± 3-sided or biconvex, smooth to rugose, apex beaked or not.

About 77 species: worldwide; 22 species (five endemic) in China.


1 Perianth bristles plumosely fringed   (2)
+ Perianth bristles retrorsely scabrous   (3)
       
2 (1) Culms terete; perianth bristles longer than nutlet; stigmas 2.   1 S. subulatus
+ Culms 3-angled to obtusely 3-angled just below inflorescence; perianth bristles ± as long as nutlet; stigmas 3.   2 S. litoralis
       
3 (1) Glumes apex emarginate and with a 1-veined costa excurrent into an awn; nutlet smooth; culms scattered; leaves developed   (4)
+ Glumes apex acute to obtuse and without an awn; nutlet mostly rugose; culms tufted (except S. lineolatus); leaves absent   (7)
       
4 (3) Involucral bracts much overtopping inflorescence, 1.5-2.5 cm; creeping rhizomes not red.   3 S. ehrenbergii
+ Involucral bracts usually shorter than inflorescence, 1.5-5.5 cm; creeping rhizomes red   (5)
       
5 (4) Culms sharply 3-angled but sometimes basally terete and only 3-angled apically; creeping rhizomes slender.   4 S. triqueter
+ Culms terete; creeping rhizomes stout   (6)
       
6 (5) Spikelets 4-7(-10) mm; glumes abaxially with reddish resinous spots; stigmas 2; nutlet plano-convex.   5 S. tabernaemontani
+ Spikelets 6-15 mm; glumes abaxially smooth; stigmas usually 3; nutlet compressed 3-sided.   6 S. lacustris
       
7 (3) Culms 3-angled (3-5-angled in S. trapezoideus and S. monocephalus), sometimes winged   (8)
+ Culms terete   (10)
       
8 (7) Perianth bristles longer than nutlet, retrorsely scabrous; nutlet compressed 3-sided.   7 S. mucronatus
+ Perianth bristles shorter than nutlet, retrorsely scabrous or not; nutlet 3-sided or biconvex   (9)
       
9 (8) Nutlet 3-sided; perianth bristles glabrous.   8 S. chuanus
+ Nutlet biconvex; perianth bristles retrorsely scabrous.   9 S. trapezoideus
       
10 (7) Plants with a creeping rhizome; culms scattered   (11)
+ Plants without a creeping rhizome; culms densely tufted   (12)
       
11 (10) Glumes oblong to elliptic, margin not ciliate; perianth bristles as long to 2 × as long as nutlet.   10 S. lineolatus
+ Glumes broadly ovate, margin ciliate; perianth bristles absent or 1-3 and very short.   11 S. monocephalus
       
12 (10) Culms 56-120 cm tall   (13)
+ Culms 15-30(-?40) cm tall   (14)
       
13 (12) Involucral bracts often deflexed; culms without transverse septa; stigmas 2.   12 S. clemensii
+ Involucral bracts erect; culms with many transverse septa; stigmas 3.   13 S. articulatus
       
14 (12) Nutlet 3-sided, distinctly rugulose; stigmas 3   (15)
+ Nutlet biconvex, obscurely rugulose or smooth; stigmas 2   (17)
       
15 (14) Perianth bristles very well developed, slightly longer than nutlet, retrorsely scabrous.   20 S. chen-moui
+ Perianth bristles very short and glabrous or absent   (16)
       
16 (15) Culms 2.5-3.5 mm thick; involucral bract 10-30 cm; perianth bristles very short, glabrous.   21 S. schoofii
+ Culms ca. 1 mm thick; involucral bract 1.5-12 cm; perianth bristles absent.   22 S. supinus
       
17 (14) Glumes oblong, 2-3 mm; nutlet 1-1.5 mm   (18)
+ Glumes broadly ovate, oblong-ovate, or elliptic, 3.5-5.5 mm; nutlet 1.6-2.5 mm   (19)
       
18 (17) Spikelets yellow, 7-20 mm; glume margin apically ciliate; nutlet obscurely rugulose.   18 S. jingmenensis
+ Spikelets brownish green, 4-7 mm; glume margin entire; nutlet subsmooth.   19 S. komarovii
       
19 (17) Glumes pale green to brownish green, oblong-ovate, 4-5.5 mm, apex acuminate; nutlet broadly ellipsoid, ca. 2.5 mm; stigmas 2; perianth bristles 4, longer than nutlet.   17 S. wallichii
+ Glumes brown to dark brown, broadly ovate to ovate, 3.5-4 mm, apex acute; nutlet broadly obovoid to obovoid, 2-2.5 mm; stigmas 3; perianth bristles 5 or 6, shorter than or as long as nutlet   (20)
       
20 (19) Culms spaced in row along rhizome; involucral bracts horizontal; glumes 3-3.2 mm.   16 S. fuscorubens
+ Culms tufted; involucral bracts ± erect; glumes to 4.5 mm   (21)
       
21 (20) Culms 0.7-1 mm thick, rigid, shiny; spikelets ovoid, 4-6 mm wide; nutlets trigonous; stigmas 3.   15 S. hotarui
+ Culms 1.5-7 mm thick, subrigid, opaque; spikelets elliptic to subcylindric, 3.5-5 mm wide; nutlets plano-convex; stigmas 2.   14 S. juncoides

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