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11. Scutellaria Linnaeus, Sp. Pl. 2: 598. 1753.
黄芩属 huang qin shu
Herbs or subshrubs, rarely shrubs, not aromatic. Leaves entire to pinnatifid. Inflorescences terminal or axillary racemes or spikes; floral leaves usually bractlike apically. Flowers axillary, opposite or sometimes alternate apically. Calyx short tubular, dorsiventrally flattened, 2-lipped; lips entire, closed and ultimately divided to base along sutures in fruit; upper lip deciduous, with a transverse, rounded, concave, scalelike scutellum (shield) or without and abaxially conspicuously saccate; lower lip persistent. Corolla 2-lipped; tube exserted, arcuate or suberect, gradually widening to throat, base bent and saccate or spurred, usually puberulent annulate outside; upper lip erect, galeate; lower lip 3-lobed, middle lobe broad, flattened, entire, lateral lobes ± joined to upper lip and sometimes spreading. Stamens 4, didynamous, anterior 2 longest, underlying upper lip; anthers close together in pairs, bearded on cell aperture; posterior pair conspicuously 2-celled, ± apically acute, anterior pair 1-celled by abortion. Style subulate, apically unequally 2-cleft. Nutlets oblate, globose, to ovoid.
About 350 species: worldwide, but only a few in tropical Africa; 98 species in China.
A very isolated genus with unsatisfactory traditional divisions. Paton (Kew Bull. 45: 399-450. 1990) has proposed a revised classification of Scutellaria but did not deal with a large
number of the Chinese taxa, and it is not possible to utilize his divisions in detail.
Key_start
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1 |
Inflorescences crowded 4-sided spikes; flowers opposite and decussate, subtended by specialized, membranous or submembranous, sessile, hoodlike bracts |
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Inflorescences 1-sided or rarely partly spiral, often poorly defined; flowers subtended by leaves or leaflike bracts. |
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(2) |
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2 (1) |
Upper lip of calyx convex, lacking a scutellum; nutlets gray-black, with hairs longer than 0.5 mm |
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98 Scutellaria kingiana |
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Upper lip of calyx with a sail-like scutellum; nutlets of various colors, glabrous or with hairs shorter than 0.5 mm. |
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(3) |
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3 (2) |
Apical flowers often ± spiral; nutlets with acuminate papillae terminating in a ring of hooks; inflorescences terminal, often ± scapose; bracts minute, herbaceous |
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1 Scutellaria discolor |
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Flowers erect, opposite; nutlets variously ornamented, mostly lacking hooked papillae; inflorescences robust and/or lateral, never scapose; bracts leaflike. |
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(4) |
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4 (3) |
Flowers in terminal racemes or spikes ± sharply differentiated from the vegetative part of the stem, mostly subtended by reduced bractlike leaves. |
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(5) |
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Flowers in axillary racemes or, if terminal, most flowers subtended by normal leaves and with a gradual transition from these to bracts. |
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(6) |
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5 (4) |
Stem leaves conspicuously petiolate, almost 2 × as long as wide, mostly obscurely dentate or rarely entire; bracts entire, short petiolate, unlike stem leaves; plants subshrubs or erect or rarely ascending herbs lacking enlarged rhizomes |
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Key 1 (see List of Keys below) |
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Stem leaves subsessile or short petiolate, more than 2 × as long as wide, dentate-serrate, crenate, to entire; bracts ± similar to stem leaves, gradually reduced toward stem apex; plants erect or ascending-erect perennial herbs with an enlarged and often woody rootstock/rhizome |
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Key 4 (see List of Keys below) |
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6 (4) |
Flowers mainly in axillary racemes or on lateral branches; plants much branched herbs |
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Key 5 (see List of Keys below) |
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Flowers axillary, secund, mostly on main axis; plants perennial, mostly erect herbs. |
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(7) |
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7 (6) |
Nutlets inconspicuously differentiated into adaxial and abaxial sides, tuberculate |
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Key 6 (see List of Keys below) |
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Nutlets conspicuously differentiated into adaxial and abaxial sides, abaxially tuberculate, adaxially spiny or smooth, with or without a membranous wing on equatorial plane. |
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(8) |
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8 (7) |
Nutlets tuberculate abaxially, adaxially smooth and conical, with an apical umbo; rhizomes with long leafless stolons at nodes; stems densely spreading pilose |
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96 Scutellaria tuberifera |
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Nutlets densely tuberculate abaxially, adaxially densely spiny around a cylindric central umbo; rhizomes without stolons; stems glabrous to sparsely pilose apically |
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97 Scutellaria guilielmii |
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List of Keys
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List of lower taxa
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