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FOC | Family List | FOC Vol. 18 | Scrophulariaceae

54. Pedicularis ser. Verticillatae Maximowicz, Bull. Acad. Imp. Sci. Saint-Pétersbourg. 32: 524. 1888.

轮叶系 lun ye xi

Ser. Amoenae Prain; ser. Plicatae Prain; ser. Pygmaea Prain.

Herbs annual or perennial, low or tall, sometimes woody. Leaves in whorls of 3 or 4, some opposite proximally, elliptic to lanceolate, pinnatisect or rarely pinnatifid. Inflorescences spicate to racemose. Calyx lobes 5 or sometimes by fusion appearing 3. Corolla usually purple, red, or white, rarely yellow; tube decurved basally; galea toothless, beakless.


1 Galea of corolla ca. 1/2 as long as lower lip.   (2)
+ Galea of corolla ca. as long as lower lip.   (9)
       
2 (1) Calyx lobes by fusion appearing 3.   (3)
+ Calyx lobes distinctly 5.   (4)
       
3 (2) Leaf segments 6--9 pairs, linear-oblong to triangular-ovate; capsule triangular-lanceolate, ca. 12 X 4 mm   149 Pedicularis holocalyx
+ Leaf segments 9 or 10 pairs, ovate to oblong; capsule obliquely ovoid, 6--7 X 4 mm   150 Pedicularis spicata
       
4 (2) Leaves pinnatifid   159 Pedicularis lineata
+ Leaves pinnatipartite to pinnatisect.   (5)
       
5 (4) Filaments glabrous.   (6)
+ 2 filaments pubescent.   (7)
       
6 (5) Roots single, not fleshy; calyx lobes triangular, ± entire or obscurely serrate   156 Pedicularis brachycrania
+ Roots usually several, fleshy; calyx lobes ovate to oblong, pinnatifid   131 Pedicularis roylei
       
7 (5) Plants with clustered fibrous roots; calyx lobes linear-lanceolate   133 Pedicularis likiangensis
+ Plants with a main root; calyx lobes wider, ± ovate or wider than long.   (8)
       
8 (7) Crown not scaly; calyx lobes unequal, posterior one triangular, entire   154 Pedicularis diffusa
+ Crown ovate scaly; calyx lobes ± equal, serrate   155 Pedicularis nyalamensis
       
9 (1) Calyx deeply cleft anteriorly.   (10)
+ Calyx not or slightly cleft anteriorly.   (18)
       
10 (9) Median or sometimes distal bracts subpalmatifid, usually with 3(--5) lobules.   (11)
+ Bracts pinnatifid or only serrate.   (12)
       
11 (10) Plants 8 cm tall or more; calyx lobes slender, 2/7--2/5 as long as calyx tube, serrulate; corolla tube decurved at middle; apex of galea not mucronulate   134 Pedicularis violascens
+ Plants barely 3 cm tall; calyx lobes triangular, short, barely as long as 1/5 calyx tube, entire; corolla tube decurved ± basally; apex of galea mucronulate   137 Pedicularis pygmaea
       
12 (10) Calyx with obscurely reticulate veins, lobes prominent.   (13)
+ Calyx with distinct reticulate veins, lobes obscure.   (16)
       
13 (12) Plants usually woody, many branched apically   136 Pedicularis chingii
+ Plants herbaceous, unbranched apically.   (14)
       
14 (13) Roots fleshy; calyx lobes distinctly lobulate   130 Pedicularis rupicola
+ Roots not fleshy; calyx lobes without lobules.   (15)
       
15 (14) Leaves 2.5--4 X as long as wide, with 6--10 pairs of segments; corolla more than 10 mm; calyx lobes grouped posteriorly; anterior filaments pubescent   135 Pedicularis verticillata
+ Leaves less than 2 X as long as wide, with 3 or 4 pairs of segments; corolla 9--10 mm; calyx lobes not grouped posteriorly; all filaments glabrous   158 Pedicularis minutilabris
       
16 (12) Proximal segments of leaf blade becoming leafletlike; corolla tube ca. 2.5 mm wide, strongly decurved basally   148 Pedicularis confluens
+ Proximal segments of leaf blade not becoming leafletlike; corolla tube 0.8--1.5 mm wide, slightly decurved basally.   (17)
       
17 (16) Leaf segments oblong-triangular; calyx 5-lobed, tube 4.5--5.5 mm, 1/2 cleft anteriorly; 2 filaments sparsely pubescent   146 Pedicularis transmorrisonensis
+ Leaf segments linear-lanceolate; calyx 3-lobed, tube ca. 3.5 mm, ca. 1/3 cleft anteriorly; 2 filaments villous   147 Pedicularis ganpinensis
       
18 (9) Plants less than 3 cm tall; calyx lobes distinctly 3-divided   143 Pedicularis minima
+ Plants more than 3 cm tall; calyx lobes not distinctly 3-lobed.   (19)
       
19 (18) Corolla tube ca. 3 X as long as galea; apex of galea ovate, acute   145 Pedicularis altifrontalis
+ Corolla tube ca. 2 X as long as galea; apex of galea rounded or truncate.   (20)
       
20 (19) Leaves pinnatifid.   (21)
+ Leaves ± pinnatisect.   (24)
       
21 (20) Lower corolla lip ciliate; calyx lobes ± triangular.   (22)
+ Lower corolla lip not ciliate; calyx lobes oblong to lanceolate.   (23)
       
22 (21) Stems usually numerous, cespitose; lobes of lower corolla lip not rounded   152 Pedicularis triangularidens
+ Stems single; lobes of lower corolla lip rounded   153 Pedicularis dolichostachya
       
23 (21) Stems rigid, usually unbranched; calyx ca. 4 mm, lobes ± serrate; plants pubescent   151 Pedicularis szetschuanica
+ Stems many branched, branches slender and weak; calyx 2--5 cm, lobes entire; plants glabrous   157 Pedicularis flaccida
       
24 (20) Median and distal bracts subpalmatifid, 3--5-lobulate, slender, fingerlike.   (25)
+ Bracts pinnatifid or slightly palmatifid, lobes short and rounded.   (26)
       
25 (24) Roots ± fleshy; leaves with linear, dentate segments; calyx lobes equal, ± entire   138 Pedicularis anthemifolia
+ Roots ± woody; leaves with lanceolate, pinnatipartite segments; calyx lobes unequal, serrate   139 Pedicularis kansuensis
       
26 (24) 2 filaments or all 4 pubescent.   (27)
+ Filaments glabrous.   (29)
       
27 (26) Bracts broadly ovate; middle lobe of lower corolla lip ca. 3.5 mm wide   144 Pedicularis tenera
+ Bracts linear-lanceolate; middle lobe of lower corolla lip ca. 2.5 mm wide.   (28)
       
28 (27) Calyx glabrous; galea not crested; lower corolla lip glabrous, apex of lobes not apiculate   140 Pedicularis glabrescens
+ Calyx long ciliate; galea narrowly crested; lower corolla lip ciliate, apex of lobes apiculate   141 Pedicularis zayuensis
       
29 (26) Calyx lobes ca. as long as calyx tube, ± pinnatisect apically, divisions slender   142 Pedicularis daucifolia
+ Calyx lobes much shorter than calyx tube, barely serrate or entire.   (30)
       
30 (29) Calyx densely whitish villous, lobes ca. 1/3 as long as calyx, unequal, ± ovate-oblong, serrate; galea 5--8 mm   131 Pedicularis roylei
+ Calyx glabrous, lobes ca. 1/5 as long as calyx, ± equal, triangular, ± entire; galea ca. 4 mm   132 Pedicularis bomiensis

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