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FOC | Family List | FOC Vol. 7 | Papaveraceae | Corydalis

10. Corydalis sect. Corydalis

实心延胡索组 shi xin yan hu suo zu

Corydalis sect. Pes-gallinaceae Irmisch; C. sect. Raphanituber Khokhryakov.

Seedlings with 1 cotyledon. Tuber solid, rounded, 0.5-3 cm in diam., base with a single fascicle of roots, apex with a tuft of small scales, from where usually (1 or)2 stems grow; each flowering stem regenerating 1 new tuber. Stems attenuate toward base, with 1 amplexicaul membranous ovate prophyll (scale leaf) at ground level, simple or with late branches from scale leaf, rarely branched from cauline leaves. Radical leaves not present in flowering individuals; cauline leaves 2 (rarely 1 or 3), alternate, petiolate, once to 4 × ternately or pinnately divided. Inflorescence a simple raceme; bracts green, large, entire to divided. Sepals usually small. Outer petals without crest, apex usually broad, obtuse to deeply emarginate; nectary obtuse to acuminate. Stigma squarish, flat, with marginal simple papillae and geminate papillae laterally and in basal corners or on basal lobes.

About 54 species: C Europe to E Asia, and N Algeria; 15 species (seven endemic) in China.

1 Nectary apically acute to acuminate   (2)
+ Nectary obtuse   (10)
2 (1) Stem straight, erect; capsule linear, or if elliptic, then flowers "bladdery."   (3)
+ Stem geniculate at nodes, ascending; capsule elliptic, with 2 rows of seeds   (6)
3 (2) Nectary extended through ca. 2/3 of spur; leaves ternate to biternate   (4)
+ Nectary extended through 1/3 or less of spur; leaves biternate to triternate   (5)
4 (3) Lower petal strongly saccate at base; capsule obovoid, ca. 1 cm.   49 C. saccata
+ Lower petal straight; capsule linear to oblong, to 2 cm.   46 C. linjiangensis
5 (3) Lower petal not or slightly saccate; capsule linear-oblong.   47 C. fumariifolia
+ Lower petal very broadly saccate; capsule elliptic; flowers "bladdery".   48 C. ussuriensis
6 (2) Inner petals 8 mm or more; dorsal crests of inner petals not or slightly extended beyond apex   (7)
+ Either inner petals shorter (to 7 mm) or dorsal appendages/crests of inner petals clearly extended beyond apex or both   (8)
7 (6) Flowers pale indigo; inner petals almost purple; lower petal base obviously saccate; capsule obovoid, ca. 1 cm.   49 C. saccata
+ Flowers blue; inner petals pale; lower petal straight, almost without sac; capsule fusiform, 1.5-2.2 cm.   50 C. humilis
8 (6) Spur sigmoidally upcurved, 11-14 mm; inner petals 9-10 mm (dorsal crest excluded).   51 C. kiautschouensis
+ Spur not or slightly curved, 6-9 mm; inner petals 5-8 mm   (9)
9 (8) Dorsal appendages of inner petals obtuse, overtopping petal apex 1/7-1/5.   52 C. repens
+ Dorsal appendages of inner petals narrowly acute, extended beyond apex 1/3 of length of petal.   53 C. watanabei
10 (1) Nectary very short, less than 1/4 as long as spur; flowers pink (Xinjiang)   (11)
+ Nectary 1/3-2/3 as long as spur; flowers blue to purple, rarely pink (E and NE China)   (12)
11 (10) Flowers 2-2.5 cm; flowers broad at apex; capsule lanceolate.   54 C. glaucescens
+ Flowers 3.5-4.5 cm; flowers narrow; capsule linear.   55 C. schanginii
12 (10) Ultimate leaflets rounded with long thin petiolules; fruit ovoid; racemes 1-8-flowered; pedicels much longer than bracts; stems geniculate, much branched   (13)
+ Petiolules short or indistinct; racemes 8-20-flowered; pedicels equaling bracts; stems usually erect   (14)
13 (12) Flowers blue or indigo, ca. 2 cm; spur 1.2-1.4 cm; nectary extended through ca. 3/4 of spur; inner petals 7-8 mm; raceme 3-8-flowered.   56 C. caudata
+ Flowers white, 1-1.2 cm; spur 5-7 mm; nectary extended through 1/3-1/2 of spur; inner petals ca. 4 mm; raceme 1-3-flowered.   57 C. humosa
14 (12) Most bracts entire; leaflets lanceolate, entire; corolla upcurved.   60 C. yanhusuo
+ Most bracts divided; leaflets lobed or dentate; corolla straight or curved   (15)
15 (14) Stems stout, simple or branched from prophyll; petioles basally not vaginate; outer petals dentate, apex retuse and mucronulate.   58 C. turtschaninovii
+ Stems weak, branched from cauline leaves; lower leaves basally vaginate; petal limbs entire and not or only slightly mucronulate.   59 C. gamosepala

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