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4. Artemisia sect. Viscidipubes Y. R. Ling, Acta Phytotax. Sin. 18: 506. 1980.

腺毛蒿组 xian mao hao zu

Herbs, perennial, rarely subshrubs; most parts usually finely arachnoid puberulent to tomentose, also with thick multicellular viscid hairs and/or gland-tipped hairs, and sessile glands. Leaves 1- or 2-pinnatipartite to pinnatisect; lobules 1-8 mm wide. Capitula ovoid or oblong-ovoid. Phyllary margins scarious, midvein green or sometimes brown. Receptacle glabrous. Marginal florets 2-15; corolla narrowly tubular or rarely narrowly conical, 2- or 3-toothed or without teeth. Disk florets 2-20(-50), bisexual; ovaries well developed; style ca. as long as or longer than corolla, branches divergent, recurved at apex.

Twenty-one species: mainly at middle and high elevations in NW and SW China, a few species in S and SE Asia; 20 species (16 endemic) in China.

Many species contain sesquiterpenoids and some monoterpenoids.

1 Middle stem leaves with 2-4 pairs of main segments   (2)
+ Middle stem leaves with 4-6 pairs of main segments   (10)
2 (1) Capitula 3-4 mm in diam., in groups of 3-6(-8) on branches of large broad panicles.   116 A. erlangshanensis
+ Capitula 1.5-3(-4) mm in diam., uniformly distributed along branches in narrow or broad panicles   (3)
3 (2) Stem and branches with many sessile blunt multicellular hairs and scattered gland-tipped hairs; middle stem leaves sessile, lobes 3-6(-8) mm wide, revolute, adaxially glandular pubescent and sparsely arachnoid pubescent, abaxial veins with hairs obviously differently colored from those elsewhere.   118 A. chingii
+ Stem and branches sparsely pubescent and glandular pubescent or not; middle stem leaves shortly petiolate, lobes (3-)10 mm wide or more, flat, adaxially uniformly glandular pubescent or on veins only or mixed with non-glandular hairs, abaxial veins with hairs same or different colors as those elsewhere   (4)
4 (3) Lobes of leaves linear or linear-lanceolate, (1-)1.5-2 × 0.3-1 cm; capitula in broad panicles   (5)
+ Lobes of leaves elliptic, oblong, or oblong-ovate, more than 2 × 1 cm; capitula in narrower panicles   (6)
5 (4) Lobes of leaves linear, 10-15(-30) × 3-6 mm.   119 A. simulans
+ Lobes of leaves linear-lanceolate or elliptic-lanceolate, 15-20 × 5-10 mm.   125 A. rosthornii
6 (4) Middle stem leaves 3- or ternately 5-partite, distal and lateral lobes subequal, not caudate, entire or few serrate at margin.   126 A. sichuanensis
+ Middle stem leaves 1- or 2-pinnatisect, or -partite, segments 2-4(or 5) pairs, distal ones longer, or not and then with broadly winged rachis, usually serrate   (7)
7 (6) Distal lobes not caudate, 2- or 3-cleft, rachis broadly winged   (8)
+ Distal lobes caudate, longer than other lobes, serrate, rachis not winged   (9)
8 (7) Leaves papery; capitula oblong or oblong-ovoid, 2.5-3(-4) mm in diam.   124 A. thellungiana
+ Leaves thickly papery; capitula ovoid, 1.5-2 mm in diam.   122 A. orientaliyunnanensis
9 (7) Leaves gland-dotted, segments 2 or 3 pairs, 1- or 2-cleft; capitula oblong or ovoid, 1.5-2(-2.5) mm in diam.   117 A. atrovirens
+ Leaves not gland-dotted, segments 3(or 4) pairs, 2- or 3-lobuled or serrate; capitula hemispheric or broadly ovoid, 2.5-3 mm in diam.   127 A. zayuensis
10 (1) Capitula (3-)4-5 mm in diam., disk florets 30-40(-50); phyllaries tomentose   (11)
+ Capitula 1.5-3(-4) mm in diam., disk florets 2-20(-25), if capitula 3-4 mm in diam. then phyllaries glabrous or sparsely puberulent   (12)
11 (10) Phyllary margins not barbate; perennial herbs, 40-120 cm tall; stems glandular pubescent.   110 A. viscida
+ Phyllary margins barbate; subshrubs, 25-50 cm tall; stems densely yellowish viscid tomentose.   111 A. vexans
12 (10) Middle stem leaves (2 or)3-pinnatisect or -partite, deeply serrate   (13)
+ Middle stem leaves 1- or 2-pinnatisect or -partite   (15)
13 (12) Uppermost flowering branches short, forming narrow panicle; phyllaries barbate at apex; capitula 3-4 mm in diam.   113 A. mattfeldii
+ Flowering branches long, forming broad, much-branched panicles; phyllaries not barbate; capitula 1-2.5 mm in diam   (14)
14 (13) Stem, branches, leaves, and phyllaries with prominently gland-tipped hairs; capitula ellipsoid or oblong, 2-2.5 mm in diam.   120 A. gyitangensis
+ Stem, branches, and leaves viscid pubescent, phyllaries glandular pubescent and eglandular pubescent or not; capitula ovoid, 1-1.5 mm in diam.   128 A. gongshanensis
15 (12) Capitula 3-4 mm in diam   (16)
+ Capitula 1.5-2.5(-3) mm in diam   (17)
16 (15) Stem and branches sparsely glandular pubescent and densely tomentose, sparsely so later; capitula in narrow panicles.   112 A. tangutica
+ Stem and branches densely glandular pubescent, not tomentose; capitula in somewhat broad panicles.   115 A. viscidissima
17 (15) Stem and branches densely viscid pubescent; leaves sparsely pubescent adaxially and densely arachnoid tomentose and white gland-dotted abaxially and sparsely glandular pubescent on abaxial veins; capitula in somewhat broad panicles; phyllaries densely tomentose and sparsely glandular pubescent.   129 A. yadongensis
+ Stem and branches densely glandular pubescent and sparsely eglandular puberulent, adaxial surface of leaves and abaxial veins glandular pubescent or also tomentose adaxially; capitula in broad panicles; phyllaries arachnoid tomentose or glabrescent   (18)
18 (17) Middle stem leaves 1-pinnatisect.   123 A. shennongjiaensis
+ Middle stem leaves 2-pinnatipartite or -pinnatisect   (19)
19 (18) Middle stem leaves 2-pinnatisect, distances between segments long, distal lobule longer than laterals, linear or linear-lanceolate, 15-25 × 1.5-2.5 mm.   114 A. occidentalisichuanensis
+ Middle stem leaves 2-pinnatipartite or nearly -sect, distances between segments shorter, distal lobule subequal to laterals, elliptic-lanceolate or ovate-elliptic, 10-15 × 3-5 mm.   121 A. myriantha

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