1 |
Filaments all free from one another form base to apex |
|
(2) |
+ |
Filaments partly or almost wholly united, either monadelphous or diadelphous |
|
(3) |
|
|
|
|
2 (1) |
Leaves pinnately foliolate |
|
Sophoreae |
+ |
Leaves digitately foliolate |
|
Podalyrieae |
|
|
|
|
3 (1) |
Anthers uniform in size and shape or nearly so, not alternately basifixed and dorsifixed |
|
(4) |
+ |
Anthers dimorphic, alternately dorsifixed and basifixed, either all equal or alternately long and short |
|
(53) |
|
|
|
|
4 (3) |
Free upper part of filaments more or less filiform,
not dilated upwards |
|
(5) |
+ |
Free upper part of all or half of the filaments dilated or expanded upwards (lower pair of leaflets
sometimes resembling stipules) |
|
(47) |
|
|
|
|
5 (4) |
Fruits not breaking up into separate joints when
mature, though sometimes filled between the seeds or divided by a longitudinal septum, sometimes 1-Seeded |
|
(6) |
+ |
Fruits jointed when ripe and transversely septate,
usually breaking up into 1-seeded segments |
|
(45) |
|
|
|
|
6 (5) |
Filaments monadelphous and partly connate into a single more or less closed tube (neither unilaterally nor bilaterally split or only partially so), sometimes the adaxial filament free at the base but connate higher up with the remainder |
|
(7) |
+ |
Filaments diadelphous, mostly the adaxial (vexillary) filaments free or partly adnate to the remainder, or if monadelphous then all more or less connate into a sheath split adaxially and more rarely also abaxially |
|
(15) |
|
|
|
|
7 (6) |
Racemes or panicles not swollen at the nodes of the rachis (at the insertion of the pedicels), or the
inflorescence fasciculate, capitate or subumbellate |
|
(8) |
+ |
Racemes with swollen (tumid) nodes at the insertion of the pedicels; bracts small or very caducous; style glabrous; calyxlobes 4 (upper 2 connate), vary rarely unequally 2-lipped |
|
Diocleae |
|
|
|
|
8 (7) |
Trees or shrubs, rarely climbers and herbs; indumentum when present not composed of biramous hairs; style glabrous or hairy; anthers neither tipped by gland nor mucronate |
|
(9) |
+ |
Herbs or shrublets with indumentum Composed mainly of biramous hairs; racemose or spikes axillary; anthers apiculate or gland tipped |
|
Indigofereae |
|
|
|
|
9 (8) |
Fruit dehiscent |
|
(10) |
+ |
Fruit indehiscent |
|
(13) |
|
|
|
|
10 (9) |
Flowers in axillary racemes or fasciculate at the older leafless nodes; leaves pinnate |
|
Robinieae |
+ |
Flowers in terminal racemes or panicles and sometimes also in the upper axils or flowers axillary |
|
(11) |
|
|
|
|
11 (10) |
Trees or shrubs, sometimes climbing |
|
Millettieae |
+ |
Shrublets or herbs, rarely annuals, not climbing (vexillum often hairy outside) |
|
(12) |
|
|
|
|
12 (11) |
Stipules more or less sagittate (perennial herbs) |
|
Galegeae |
+ |
Stipules not sagittate or absent (shrublets or herbs) |
|
Tephrosieae |
|
|
|
|
13 (9) |
Fruit not semi-orbicular or circinate; usually more than 2-seeded |
|
(14) |
+ |
Fruit semi-orbicular or circinate, 2-1-seeded; herbs or shrublets, rarely spiny |
|
Hedysareae |
|
|
|
|
14 (13) |
Calyx mostly cyathiform, truncate with small teeth; vexillum often silky outside; trees or tall scrambling shrubs |
|
Lonchocarpeae |
+ |
Calyx not cyathiform; shrublets or herbs; flowers mostly capitate or umbellate |
|
Loteae |
|
|
|
|
15 (6) |
Leaves simple (not unifoliolate), neither reduced to scales nor phyllodes; herbs with biramous hairs and apiculate stamens |
|
Indigofereae |
+ |
Leaves compound |
|
(16) |
|
|
|
|
16 (15) |
Leaves digitately foliolate, sometimes unifoliolate |
|
(17) |
+ |
Leaves pinnately trifoliolate or pinnate |
|
(15) |
|
|
|
|
17 (16) |
Leaves neither gland-dotted nor pellucid-punctate below |
|
(18) |
+ |
Leaves glandular or pellucid-punctate below |
|
(24) |
|
|
|
|
18 (17) |
Fruit not longitudinally septate i.e, with out an intrusion of the seed-bearing suture |
|
(19) |
+ |
Fruit longitudinally septate or partly so by the intrusion of the seed-bearing suture; stipels absent |
|
Astragaleae |
|
|
|
|
19 (18) |
Shrubs or herbs with indumentum (when present) composed of simple hairs; anthers not apiculate |
|
(20) |
+ |
Herbs or shrublets with indumentum (when present) composed of biramous hairs, anthers apiculate |
|
Indigofereae |
|
|
|
|
20 (19) |
Leaflets not toothed, mostly shrubs, rarely herbs |
|
(21) |
+ |
Leaflets more or less toothed, the nerves continued to the margin; mostly herbs with trifoliolate leaves; stipules often adnate to the petiole, stipels absent |
|
Trifolieae |
|
|
|
|
21 (20) |
Ovary with 1 ovule; fruit 1-seeded |
|
Lespedezeae |
+ |
Ovary with more than 1 ovule |
|
(22) |
|
|
|
|
22 (21) |
Stipels absent; mostly shrubs or shrublets; flowers in terminal (rarely leaf-opposed) spikes, racemes or heads, rarely axillary |
|
Lotononideae |
+ |
Stipels present; mostly herbs, often climbing |
|
(23) |
|
|
|
|
23 (22) |
Fruit jointed but dehiscing the full length |
|
Desmodieae |
+ |
Fruits not jointed, though sometimes filled between the seeds |
|
Glycineae |
|
|
|
|
24 (17) |
Fruits dehiscing by valves, 2 or more seeded |
|
Cajaneae |
+ |
Fruits indehiscent, usually 1-seeded |
|
Psoraleeae |
|
|
|
|
25 |
Leaves gland-dotted or pellucid-punctate below; ovules few (4-1) |
|
(26) |
+ |
Leaves neither gland dotted nor pellucid-punctate; ovules usually more than 4, rarely 2 or 1 |
|
(29) |
|
|
|
|
26 (25) |
Fruits dehiscent; seeds mostly 2 or more, rarely 1 |
|
(27) |
+ |
Fruits usually indehiscent and nearly always 1-seeded, bracteoles absent or minute and caducous; anthers versatile; shrubs and herbs |
|
Psoraleeae |
|
|
|
|
27 (26) |
Bracteoles mostly present; leaves impair-pinnate or 3-or 1-foliolate; vexillary filament free or free at the base |
|
Millettieae |
+ |
Bracteoles absent; leaves 3-foliolate, rarely 1-foliolate |
|
Cajaneae |
|
|
|
|
28 |
Fruit dehiscent by a valve or valves (sometimes only
at the apex) |
|
(29) |
+ |
Fruit indehiscent, sometimes inflated and bladder-
like |
|
(68) |
|
|
|
|
29 (28) |
Petals very unequal in length, either the vexillum or the keel petals much longer than the others or each other; stipels usually present, sometimes gland-like; seeds estrophiolate |
|
Erythrineae |
+ |
Petals more or less equal in length (or at any rate the keel-petals not much longer than the others; though sometimes beaked and spirally twisted |
|
(30) |
|
|
|
|
30 (29) |
Indumentum (when present) usually composed, at least in part, of biramous hairs; anthers apiculate; stipels mostly absent |
|
Indigofereae |
+ |
Indumentum not as above, anthers not apiculate |
|
(31) |
|
|
|
|
31 (30) |
Rachis of the inflorescence thickened (nodose) at the insertion of the pedicels; leaves mostly stipellate; vexillary filament usually quite free, radicle inflexed; style bearded along the inner face; keel-petals often spirally twisted |
|
Phaseoleae |
+ |
Rachis of the inflorescence not thickened at the insertion of the pedicels, or the flower fasciculate,
capitate or axillary |
|
(32) |
|
|
|
|
32 (31) |
Rachis of the leaves transformed at the apex into a tendril or barren tip or bristle; stipels absent |
|
(33) |
+ |
Rachis of the leaves not ending in a tendril or barren tip |
|
(34) |
|
|
|
|
33 (32) |
Stamens 10; flowers solitary or inflorescence racemose in the leaf axils; claw of the vexillum
free from the staminal sheath |
|
Vicieae |
+ |
Stamens 9, the vexillary absent; flowers in terminal or subterminal racemes; claw of the vexillum more or less adnate to the staminal sheath |
|
Abreae |
|
|
|
|
34 (32) |
Fruits longitudinally septate within by the intrusion of the seed-bearing suture, often inflated; anther-loculi mostly separate at the apex; style mostly glabrous |
|
Astragaleae |
+ |
Fruit not longitudinally septate as described above, sometimes transversely septate but not breaking up into separate joints |
|
(35) |
|
|
|
|
35 (34) |
Fruit inflated and bladder-like, sometimes dehiscent only at the apex; style bearded along the adaxial side or near the apex; plants not glandular |
|
Coluteae |
+ |
Fruits not inflated, mostly compressed, though sometimes turgid around the seeds |
|
(36) |
|
|
|
|
36 (35) |
Stipules when present not adnate to the petiole or only slightly so; leaflets entire |
|
(37) |
+ |
Stipules mostly adnate to the petiole; fruits not
septate; leaflets often toothed; mostly herbs |
|
Trifolieae |
|
|
|
|
37 (36) |
Ovary with more than 1 ovule |
|
(38) |
+ |
Ovary with 1 ovule and fruit 1-seeded, enclosed by the persistent calyx; leaves pinnately 3-foliolate or 1-foliolate |
|
Lespedezeae |
|
|
|
|
38 (37) |
Style glabrous |
|
(39) |
+ |
Style bearded |
|
(43) |
|
|
|
|
39 (38) |
Racemes axillary or at the older leafless nodes |
|
(40) |
+ |
Racemes and panicles terminal or leaf opposed |
|
(42) |
|
|
|
|
40 (39) |
Hard-wooded trees and shrubs (seeds estrophiolate;
flowers in axillary racemes or fasciculate at the older nodes) |
|
Robinieae |
+ |
Herbs and herbaceous climbers or creepers, sometimes subshrubby |
|
(41) |
|
|
|
|
41 (40) |
Upright herbs, rarely tall and subshrubby; fruit
mostly filled between the seeds |
|
Sesbanieae |
+ |
Climbing herbs, rarely shrubs or trees; fruits usually not filled between the seeds |
|
Glycineae |
|
|
|
|
42 (39) |
Hard-wooded trees or climbing shrubs |
|
Millettieae |
+ |
Herbs, sometimes climbing or shrublets; stipels absent |
|
Tephrosieae |
|
|
|
|
43 (38) |
Hard-wooded trees and shrubs, rarely climbing shrubs |
|
Robinieae |
+ |
Herbs, sometimes tall or subshrubby, or rigid under-shrubs |
|
(44) |
|
|
|
|
44 (43) |
Racemes axillary or rarely flowers single or few in the leaf-axils, not leafy at the base |
|
Sesbanieae |
+ |
Racemes terminal or leaf-opposed or in the upper leaf-axils, often leafy at the base |
|
Tephrosieae |
|
|
|
|
45 (5) |
Leaves without stipels |
|
(46) |
+ |
Leaves with stipels (adaxial filaments free or more or less connate into a closed tube) |
|
Desmodieae |
|
|
|
|
46 (45) |
Keel-petals often obliquely truncate at the apex,
wings short or very small, rarely equalling the keel
Petals; filaments connate into a single sheath split
adaxially, or vexillary filaments free |
|
Hedysareae |
+ |
Keel petals obtuse or beaked, incurved; wings often Transversely plicate; filaments sometimes connate into 2 lateral bundles |
|
Aeschynomeneae |
|
|
|
|
47 (4) |
Fruits 2-valved or indehiscent, sometimes transversely septate but not breaking up into separate joints |
|
(48) |
+ |
Fruits jointed and breaking up into separate segments, these indehiscent or rarely dehiscing
by 2 valves (stipules absent, flowers not precocious) |
|
Coronilleae |
|
|
|
|
48 (47) |
Leaf-rachis not ending in a bristle or tendril |
|
(49) |
+ |
Leaf-rachis ending in a bristle or tendril; flowers
solitary or racemose in the leaf-axils |
|
Vicieae |
|
|
|
|
49 (48) |
Claws of the petals not adnate to the staminal sheath or tube |
|
(50) |
+ |
Claws of some or all of the petals partly adnate to the staminal sheath; flowers in heads or dense spikes; leaflets usually toothed, the lateral nerves
extended to the teeth or margin |
|
Trifolieae |
|
|
|
|
50 (49) |
Stamens united into an open sheath |
|
(51) |
+ |
Stamens united into a closed tube |
|
(52) |
|
|
|
|
51 (50) |
Stipules more or less free form the petiole; leaflets entire, veins not running into the margin; lowermost pair often resembling stipules |
|
Loteae |
+ |
Stipules mostly partly adnate to the petiole; veins of the leaves extended to the usually toothed margin |
|
Trifolieae |
|
|
|
|
52 (50) |
Subarborescent shrubs, silky pubescent, not glandular; leaves imparipinnate; stipules small, free; racemes terminal |
|
Tephrosieae |
+ |
Mostly perennial or annual herbs clothed with glandular hairs; leaves pinnately trifoliolate,
rarely 1-foliolate or 2-3 pairs; stipules mostly
herbaceous, adnate to the petiole |
|
Ononideae |
|
|
|
|
53 (3) |
Upper part of filaments swollen or expanded; perennial or annual herbs or shrublets, often spiny and glandular |
|
Ononideae |
+ |
Upper part of filaments neither swollen nor expanded |
|
(54) |
|
|
|
|
54 (53) |
Fruits not transversely septate or if so then not
breaking up into separate joints |
|
(55) |
+ |
Fruits transversely septate and breaking up into 1-seeded joints (sometimes some fruits reduced to 1 joint) leaves few-foliolate |
|
(67) |
|
|
|
|
55 (54) |
Leaves not glandular-punctate; anthers markedly
unequal; fruits usually more than 1-seeded |
|
Psoralea |
+ |
Leaves glandular-punctate; anthers nearly uniform;
fruits usually indehiscent and mostly 1-seeded |
|
Psoraleeae |
|
|
|
|
56 |
Leaves pinnately 3-foliolate, usually with stipels; stems often twining or climbing, sometimes erect or prostrate |
|
(57) |
+ |
Leaves digitately 3-17-foliolate, 1-foliolate, simple or reduced to scales; stipels absent; stems not twining or climbing |
|
(58) |
|
|
|
|
57 (56) |
Vexillary filaments free form the others or nearly so; petals very unequal in length |
|
Erythrineae |
+ |
Vexillary filament connate with the others. Climbing herbs; leaves 3-foliolate; stipules small; calyx scarcely 2-lipped |
|
Glycineae |
|
|
|
|
58 (56) |
Filaments in a sheath open at base or split adaxially |
|
(59) |
+ |
Filaments in a closed tube |
|
(61) |
|
|
|
|
59 (58) |
Style glabrous; keel usually obtuse or rounded;
vexillum without callus-like appendages at the base |
|
(60) |
+ |
Style ciliate; keel beaded; vexillum with paired
Callus-like appendages at the base |
|
Crotalaria |
|
|
|
|
60 (59) |
Calyx with lobes subequal or with the lateral
sinuses shallowest, rarely (in South Africa) with
a 3-fid lower lip |
|
Lotononideae |
+ |
Calyx with 3 lower lobes in a 3-fid lip |
|
Lupineae |
|
|
|
|
61 (58) |
Leaves 1-3-foliolate, simple or reduced to scales
(e.g. Spartium); keel sharply beaked (occasionally
apiculate) |
|
(62) |
+ |
Leaves 5-11 (-17)-foliolate, rarely (in America) 1-3-
foliolate, with stipules partly adnate to the petiole; keel sharply beaded, often enveloped by broad, apically coherent wings |
|
Lupineae |
|
|
|
|
62 (61) |
Calyx usually much shorter than keel-petals or with (1-) 2 short lips; stipules small from a persistent leaf-base or lacking; leaves various, often simple or reduced |
|
Genisteae |
+ |
Calyx nearly as long as the keel-petals, deeply 2-lipped; stipules usually form the branch, often expanded, sometimes adnate to the petiole; leaves 3-foliolate |
|
Lupineae |
|
|
|
|
63 |
Style glabrous |
|
(64) |
+ |
Style bearded lengthwise or ciliate on the adaxial side; fruit usually turgid or inflated |
|
Crotalarieae |
|
|
|
|
64 (63) |
Leaves pinnately foliolate; stipels mostly present; mostly climbers, the fruits sometimes with stinging hairs |
|
(65) |
+ |
Leaves digitately 3- or rarely 1-foliolate, sometimes sessile; mostly erect shrubs or shrublets; the fruit without stinging hairs |
|
(66) |
|
|
|
|
65 (64) |
Vexillary filaments free form the others or nearly so; petals very unequal in length |
|
Erythrineae |
+ |
Vexillary filaments connate with the others. Climbing herbs; leaves 3-foliolate; stipules small; calyx scarcely 2-lipped |
|
Glycineae |
|
|
|
|
66 (64) |
Calyx-lobes or teeth at most only slightly longer than the tube |
|
Lotononideae |
+ |
Calyx-lobes or lips much longer than the tube |
|
Lupineae |
|
|
|
|
67 (54) |
Filaments all connate into a closed tube; flowers spicate, capitate or rarely subracemose, terminal or axillary; fruits maturing above or below the ground |
|
Stylosantheae |
+ |
Filaments all connate into a sheath split adaxially or into 2 lateral bundles, rarely the vexillary filament free; flowers in axillary racemose rarely in axillary fascicles or subcorymbose; fruits maturing above the ground |
|
Aeschynomeneae |
|
|
|
|
68 (28) |
Fruit inflated and bladder-like or inflated on each face; stipels absent |
|
(69) |
+ |
Fruit neither inflated nor bladder-like, sometimes winged |
|
(70) |
|
|
|
|
69 (68) |
Fruit not longitudinally septate; style bearded along the adaxial (vexillary) side or near the apex; shrubs or herbs; leaves pinnate or 1-foliolate |
|
Colutea |
+ |
Fruit longitudinally septate or partly so by the intrusion of the seed-bearing suture; style glabrous |
|
Astragaleae |
|
|
|
|
70 (68) |
Leaflets (mostly) alternate, rarely only 1 leaflet.
Anthers small, didymous, not versatile, loculi erect, opening by short, terminal slit or divergent
at the base with a slit lengthwise |
|
Dalbergieae |
+ |
Leaflets opposite or a few at the same time alternate, or leaflets 3 |
|
(71) |
|
|
|
|
71 (70) |
Leaf pinnately trifoliolate or unifoliolate (ovary
with 1 ovule) |
|
Lespedezeae |
+ |
Leaf imparipinnate; vexillum often silky outside;
ovary with 2 or more ovules |
|
(72) |
|
|
|
|
72 (71) |
Trees or tall scrambling shrubs; flowers in racemes
or panicles |
|
Lonchocarpeae |
+ |
Shrublets or herbs, flowers capitate or umbellate |
|
Loteae |